Olethreutini

Agassiz, David J. L. & Aarvik, Leif, 2014, New Tortricidae (Lepidoptera) from East Africa with an account of the tortricid fauna of acacia in the Kenyan Rift Valley, Zootaxa 3861 (4), pp. 369-397 : 374-382

Olethreutini

Paraeccopsis Aarvik, 2004

The genus Paraeccopsis was established to accommodate Argyroploce insellata Meyrick, 1920 ( Aarvik 2004) . Like many other tropical species originally placed in Argyroploce by Meyrick, Paraeccopsis insellata is only distantly related to the type species of that genus, the Palaearctic Argyroploce arbutella (Linnaeus, 1758) . Razowski (2008) transferred additional African species from Argyroploce to Paraeccopsis . The discovery of a new species of Paraeccopsis feeding on acacia prompted us to study more material of the genus which led to the discovery of five additional new species. In order to facilitate the identification of the newly discovered acacia feeder and its relatives, we present figures and diagnoses of all species available to us.

Paraeccopsis variegana Agassiz & Aarvik , sp. n. ( Figs. 5–10 , 34 , 45 )

Type material. Holotype ♂, KENYA: Rift Valley, L. Naivasha, 1900 m, 0˚47’S, 36˚24’E, Acacia xanthophloea , em: 17.xi.2003, D.J.L. Agassiz ( NMK). Paratypes (18). Same data as holotype, em: 12.xi.–8.xii.2003. 5♂, 4♀ paratypes in NMK, 5♂, 4♀ in BMNH, including genitalia slide 2009.020 L. Aarvik.

Additional specimens examined. Fourteen specimens bred from A. xanthophloea ¸ K E N YA: R i f t Val l ey Province: 6 from Lake Naivasha, 2 from Rumuruti, 1 from Gatamayo Forest, 1 from Rongai, 2 from lake Nakuru Park, 1 from Malewa ( BMNH); (Lake) Naivasha, 2003, 2007; 2 bred from A. tortilis, Lake Bogoria & Logumgum, 2007 ; specimens taken at light: Kenya: Naivasha 22, 1999–2011; 2 from Rongai, 2000; 1 from Rumuruti, 1999; 2 from Lake Nakuru, 1999, 1 from Soysambu Estate, 2005; 1 from Sweetwaters, Nanyuki, 2000; Central Province: 1 from Gatamayo Forest, 1999; Coast Province: 2 from Rukinga Reserve, 2011. UGANDA: 1 from Kampala, 2000 ( DJLA). TANZANIA: Arumeru Distr.: Usa River 1170 m, 1♂, 4.viii.1991, L. Aarvik, genitalia slide L. Aarvik 2796; same locality, 1♀, 10.viii.1991, L. Aarvik, genitalia slide L. Aarvik 2797 ( NHMO).

Description. Adult ( Figs. 5–10 ). Head: Vertex and appendages pale fuscous, labial palpus tufted, segment 3 short, antenna faintly annulate. Thorax: Dorsum pale fuscous, sometimes with darker or reddish scales. Wingspan 12–15 mm. Forewing very variable, pale fuscous with areas of dark fuscous or reddish; basal blotch usually darker, often a darker triangular mark on middle of costa, termen often darker. Hindwing fuscous, a single fuscous cilia line. Abdomen: Male genitalia ( Fig. 34 ). Uncus basally broad, triangular, apically bilobed, with strong spines; socii shaped as slender lobes; valva with medial excavation in addition to the normal basal excavation, cucullus short, parallel-sided; valvae asymmetrical as the left valva has a smaller medial excavation, dorsal edge of medial excavation strongly sclerotised, with cluster of long spines; phallus long and sinuous, no carinae or cornuti.

Female genitalia ( Fig. 45 ). Strongly asymmetrical; posterior edge of sternite 7 with wide u-shaped excavation; sterigma a curved tube, sclerotised on both sides, posteriorly with two lateral large plates; ductus bursae tubular, posterior half sclerotised, anterior half membranous; ductus seminalis inserted at transition of sclerotised and membranous part; signum a round sclerite with 2–4 anteriorly directed projections of dissimilar size.

Diagnosis. Due to considerable external variation it is difficult or impossible to distinguish this species superficially from other species of the genus. The male genitalia ( Fig. 34 ) differ from those of other Paraeccopsis species mainly by the short cucullus. The female genitalia ( Fig. 45 ) are characterized by its strongly curved sterigma.

Distribution. Kenya, Uganda and Tanzania.

Biology. The larva feeds among spun leaves on Acacia xanthophloea .

Etymology. The specific epithet refers to the remarkable external variation.

Paraeccopsis exhilarata ( Meyrick, 1918) ( Figs. 11 , 35 , 46 )

Argyroploce exhilarata Meyrick 1918: 50 ; Razowski & Krüger 2007: 119, fig. 63. Paraeccopsis exhilarata: Razowski 2008: 62 .

Argyroploce acroplecta Meyrick 1921: 60 ; Razowski & Krüger 2007: 118, figs 58, 198. syn. nov. Paraeccopsis acroplecta: Razowski 2008: 62 .

Polychrosis eoplecta Meyrick 1925: 142 ; Razowski & Krüger 2007: 120, figs. 71, 284. Olethreutes eoplecta: Vári et al. 2002: 63 [synonymised with exhilarata ] Paraeccopsis eoplecta: Razowski 2008: 62 .

Material examined. [Rep. S. Africa] Blaawkraantz R. & Tugela R. Junc., 2500 ft., Natal, 1♂, xi.1896, Marshall leg.; Estcourt, 4000 ft., Weenen Natal, 1♀, i.1897 Marshall leg., genitalia slide ♀ BM 32546; Kimbolton Estcourt Weenen, Natal, 1♂, 5♀, 1885, Htchsn. leg., genitalia slide ♂ BM 32545; 2♂, 3♀, same locality, 1892, genitalia slide ♀ BM 32547, genitalia slide ♂ BM 32548 ( BMNH).

Diagnosis. In P. exhilarata the forewing reddish ochreous with brown markings more or less suffused with black scales ( Fig. 11 ). P. deltophora has a paler ground colour and more distinct markings. The pattern is similar in the two species. In the male genitalia of P. exhilarata ( Fig. 35 ) the valva is slender, and the length of cucullus about half the length of the valva. The cucullus is nearly as long in P. windhoeca Razowski, 2008 , but this species has the dorsal tuft of bristles in a more proximal position. Female genitalia ( Fig. 46 ) are very close to those of P. deltophora ( Meyrick, 1921) , and it is perhaps not always possible to separate them. The triangular sterigma posterior to ostium is slightly longer than in P. deltophora .

Distribution. Republic of South Africa and Zimbabwe. The type specimens of Argyroploce acroplecta and Polychrosis eoplecta originated from Zimbabwe; the type of Argyroploce exhilarata from the Republic of South Africa ( Razowski & Krüger 2007).

Remarks. The types of the three nominal species involved were figured by Razowski & Krüger (2007). Externally the three are virtually identical, and the synonymy of eoplecta and exhilarata was suggested by these authors. However, the synonymy of eoplecta and exhilarata had already been proposed by Vári et al. (2002). Based on the external similarity we also synonymise acroplecta . Each of the three names is based on a single type specimen; the exhilarata type lacks the abdomen, the acroplecta type is a male, and the eoplecta type is a female. All three nominal species were transferred to Paraeccopsis by Razowski (2008).

Paraeccopsis windhoeca Razowski, 2008 ( Figs. 12 , 36 )

Paraeccopsis windhoeca Razowski, 2008: 62 , figs. 4, 8.

Material examined. 1♂, S.W. Africa (W49) Rietfontein, 23 mls. SW Grootfontein, 3.iv.1972, S. Afr. Exp. B.M. 1972, genitalia slide BM 32551 ( BMNH).

Diagnosis. ( Fig. 12 ) Externally, P. windhoeca somewhat resembles P. exhilarata , but the former is darker and deeper red, with less distinct markings. In the male genitalia of P. windhoeca ( Fig. 36 ), the cucullus is rather long, but shorter than that of P. exhilarata , and the tuft of bristles dorsally on the right valva has a proximal position. For differences from P. tanzanica , sp. n., see that species. The female is not known.

Distribution. Namibia.

Remarks. Paraeccopsis windhoeca was described from a single male from Windhoek, Namibia ( Razowski 2008). The specimen figured in the present paper was discovered among unidentified Tortricidae in BMNH, and it is the second known specimen.

Paraeccopis tanzanica Aarvik , sp. n. ( Figs. 13 , 37 )

Type material. Holotype ♂, TANZANIA: Mbeya Distr.: Utengule Lodge, 26.xi.2005, L. Aarvik, M. Fibiger, A. Kingston, genitalia slide NHMO 2211 ( NHMO).

Description. Adult. Male ( Fig. 13 ). Head: Vertex and antenna light greyish brown; palps missing. Thorax: Light greyish brown; hind-legs light ochreous; fore-legs suffused with brown, mid-legs ochreous. Wingspan 15.5 mm. Forewing pale brown, suffused with brown; median fascia short, curved, only expressed near costa, together with black costal dot at 2/3 forming conspicuous blackish triangle on costa. Cilia beige, with single cilia line. Hindwing pale brownish grey with darker suffusion on veins, cilia pale beige with single cilia line. Abdomen: Male genitalia ( Fig. 37 ). Uncus basally broad, triangular, apically club-shaped, with spines; socii shaped as slender lobes; valva without medial excavation developed as in some other Paraeccopsis species, medially with strongly sclerotised ridge, armed with four large spikes, the ridge is more developed in the right valva; valva slightly sinuous, more so than in congeners; cucullus slightly shorter than half of valva, parallel-sided; phallus long and sinuous, no carinae or cornuti.

Female. Not known.

Diagnosis. Judging from the male genitalia, this species is closest to P. windhoeca . It differs from that species by a broader uncus and by the presence of four large spines on the valva, whereas P. windhoeca has more numerous and less strong spines. The valva in P. windhoeca has a longer cucullus.

Distribution. Tanzania.

Etymology. Named after Tanzania the country where the type specimen was found.

Paraeccopsis addis Aarvik , sp. n. ( Figs. 14 , 38 , 47 )

Type material. Holotype ♂, ETHIOPIA: Addis Ababa, 4.ix.1980, Dr. Angenstein, genitalia slide L. Aarvik 2012.009 ( ZMHU). Paratypes (2). Same data as holotype, 1♂, 14.ix.1980, 1♀, 26.v.1983, genitalia slide L. Aarvik 2013.041 ( ZMHU).

Description. Adult. Male ( Fig. 14 ). Head: Vertex pale brown; antenna light brown, short ciliate; labial palpus with internal surface light brown, external surface dark brown, 1.5 times diameter of eye. Thorax: Pale brown, posterior tuft blackish brown. Legs pale ochreous-brown, external surface of fore and mid-legs with dark brown suffusion, making the light ground colour visible as rings. Wingspan 16 mm. Forewing beige; sub-basal fascia blackish brown, forming dark patch in dorsal half, in costal half obsolete; a prominent triangular blotch present on costa above white discal spot; discal spot edged with blackish below; triangular area along termen from apex to 1/3 from tornus suffused with black and brown. Cilia grey, basally brown, at tornus cream-coloured. Hindwing greyish brown, veins darker; cilia dirty white, with single dark cilia line. Abdomen: Genitalia ( Fig. 38 ). Uncus basally broad, triangular, apically slightly bilobate, with long spines; socii shaped as slender lobes; valva with medial excavation in addition to the normal basal excavation, excavation of left valva less distinct, dorsal edge of excavation forming large plate on right valva, armed with cluster of long spines and setae; cucullus less than half the length of the valva, parallel-sided; phallus long and sinuous, no carinae or cornuti.

Female. Head and thorax: Similar to male. Abdomen: Genitalia ( Fig. 47 ). Strongly asymmetrical; posterior edge of sternite 7 with wide u-shaped excavation; sterigma a curved tube, posteriorly with two lateral large plates; posterior edge of segment 7 with curved, strongly sclerotised rims, one on each side; ductus bursae tubular, posterior half sclerotised, anterior half membranous; ductus seminalis inserted at transition of sclerotised and membranous part; signum a round sclerite with 2 anteriorly directed projections of dissimilar size.

Variation. The description is based on the holotype. The male paratype differs by presence of reddish brown suffusion on the forewing, particularly below the coastal triangular blotch.

Diagnosis. The male genitalia ( Fig. 38 ) differ from those of other Paraeccopsis species by the presence of a large plate in the middle of the right valva. The female genitalia ( Fig. 47 ) resemble those of P. variegana , sp. n., but differ by having the curved sterigma evenly sclerotised (not in the form of two lateral bands), and by the presence of strongly sclerotised edges posteriorly on segment 7. Externally, the new species is probably indistinguishable from P. turi , sp. n. (see below).

Distribution. Ethiopia.

Etymology. Named after the capital of Ethiopia, Addis Ababa, where the type specimens were collected.

Paraeccopsis turi Aarvik , sp. n. ( Figs. 15 , 39 , 48 )

Type material. Holotype ♂, KENYA: Rift Valley Prov.: Turi , 4.ii.2000, D.J.L. Agassiz., genitalia slide L. Aarvik 2012.007 ( BMNH). Paratypes (3). Same data as holotype, 1♂, 13.vi.2000, genitalia slide L. Aarvik 2013.022; 1♀, 12.ii.1999, genitalia slide L. Aarvik 2012.013 ( BMNH); 1♀, Rift Valley Prov., Mt. Elgon Nat. Park, Chorlim gate, Rongai camp, 2206 m, 01˚01’51,7’’N, 34˚46’40,8’’E, 21.xi.2006, L.O. Hansen & K. Sund, genitalia slide NHMO 2245 ( NHMO).

Description. Adult. Male ( Fig. 15 ). Head: Vertex pale brown; antenna light brown, short ciliate; labial palpus with internal surface light brown, external surface dark brown, 1.5 times diameter of eye. Thorax: Brown, posterior tuft with whitish scales in middle. Legs pale ochreous-brown, external surface of fore and mid-legs with dark brown suffusion, making the light ground colour visible as rings. Wingspan 17.5 mm. Forewing ochreous brown; sub-basal fascia blackish brown, forming dark patch in dorsal half, in costal half obsolete; a prominent triangular blotch present on costa above white discal spot; discal spot edged with blackish below, prolonged as pale band to costa just before apex; tornal marking yellowish white. Cilia grey, basally brown, at tornus cream-coloured. Hindwing greyish brown, veins darker; cilia dirty white, with single dark cilia line. Abdomen: Genitalia ( Fig. 39 ). Uncus basally broad, triangular, apically parallel-sided, slightly indented on distal edge, spiny; socii shaped as slender lobes; valva with medial oval excavation in addition to the normal basal excavation; edge of medial excavation sclerotised, in particular in right valva where the dorsal edge is widened to a plate carrying numerous long spines; left valva with a few spines in the same position; cucullus relatively short, slightly more than one third of length of valva, apically rounded, gradually narrowing towards apex; phallus long and sinuous, no carinae or cornuti.

Female. Head and thorax: Differing from male mainly by having thorax and forewing suffused with rufous. Abdomen: Genitalia ( Fig. 48 ). Posterior edge of sternite 7 with wide u-shaped excavation; anterior edge of ostium forming well defined triangle; sterigma broad and short, divided into two gradually narrowing bands that diverges towards the bases of the apophyses anteriores; ductus bursae tubular, posterior half sclerotised, anterior half membranous, sclerotised part rather broad; ductus seminalis inserted in membranous part just before sclerotised part; signum an irregular sclerite with 2 anteriorly directed projections of similar size.

Diagnosis. This species is closely related to Paraeccopsis deltophora ( Meyrick, 1921) . In the male genitalia ( Fig. 39 ), it differs by the larger uncus and longer cucullus. The female genitalia ( Fig. 48 ) are characterised by their short and broad ostium/sterigma complex. Paraeccopsis addis , sp. n., known from Ethiopia, is externally similar to P. turi , sp. n., but there are conspicuous differences in the genitalia of both sexes.

Distribution. Kenya.

Etymology. Named after the type locality, Turi , Kenya, where three of four specimens were collected.

Paraeccopis botswanae Aarvik , sp. n. ( Figs. 16 , 40 )

Type material. Holotype ♂, BOTSWANA SE, Mashatu: Tuli Game Reserve, Limpopo River, 530 m, 22˚12.644’S, 29˚08.402’E, 14.v.2004, A.J. Kingston, genitalia slide NHMO 2404 ( NHMO).

Description. Adult,( Fig. 16 ). Male. Head: Beige; antenna light brown, short ciliate; labial palpus beige, external surface with some brown scales, 1.5 times diameter of eye. Thorax: Pale brown, with dark grey transverse band; crest brownish red. Fore and mid-legs beige, tarsi and fore tibia with brown rings, hind leg cream coloured with faintly ringed tarsi. Wingspan 13.0 mm. Forewing ochreous; subbasal and medial fasciae brown, connected in dorsal half, forming large trapezoid patch covering most of wing, interspersed with blackish striae; a dark macula in terminal area connected with termen in middle; white discal spot small; costal strigulae weak. Cilia beige, with three grey streaks in apical area and one broad in middle. Hindwing grey, becoming lighter towards base, veins darker; cilia dirty white, with single dark cilia line. Abdomen: Genitalia ( Fig. 40 ). Uncus basally broad, triangular, spiny; socii shaped as slender lobes; valva with medial oval excavation in addition to the normal basal excavation; edge of medial excavation sclerotised, in particular in the right valva where the dorsal edge is widened to a plate carrying numerous long spines; left valva with a few spines at the distal edge of the medial excavation; cucullus short, about one third of length of valva, apically rounded, gradually narrowing towards apex; phallus long and sinuous, no carinae or cornuti.

Female. Not known.

Diagnosis. Paraeccopsis botswanae closely resembles some forms of P. variegana , sp. n. ( Fig. 5 ), with which it was initially confused. However, the male genitalia are close to those of P. t u r i, sp. n. The genitalia of P. botswanae differ in having a shorter uncus and shorter cucullus, and by having fewer spines at the distal edge of the medial excavation of the left valva. The shape of the cucullus is very similar to that of P. deltophora ( Fig. 41 ), but the latter species differs by the much smaller uncus.

Distribution. Botswana.

Etymology. Named after the country Botswana, where the type specimen was collected.

Paraeccopsis deltophora ( Meyrick, 1921) , comb. n. ( Figs. 17 , 41 , 49 )

Polychrosis deltophora Meyrick, 1921: 55 ; Razowski & Krüger 2007: 120, fig. 70.

Material examined. [Rep. S. Africa] Kimbolton Estcourt Weenen, Natal, 1♂, 2♀♀, 1892 Htchsn., genitalia slide ♂ BM 32549, ♀ BM 32550; Umtata Transkei Cape Colony 1♂, 1899, Barrett ( BMNH).

Diagnosis. In P. deltophora the forewing ground is colour light ochreous ( Fig. 17 ), with the same general pattern as P. addis sp. n., P. t u r i sp. n. and P. insellata (Meyrick) , consisting of a subbasal, oblique blackish patch on dorsum and a similarly coloured triangle on costa; the outer edge of the costal triangle touching distinct white discal spot. The male genitalia of P. deltophora ( Fig. 41 ) has a smaller uncus than all other species except P. insellata , and a relatively short valva, with a short cucullus armed with strong spines. P. insellata has longer cucullus. The female genitalia of P. deltophora ( Fig. 49 ) resemble those of P. exhilarata , but the sterigma posterior to the ostium appears to be shorter in P. deltophora than in P. exhilarata .

Distribution. Mozambique and Republic of South Africa.

Remarks. Meyrick (1921) described Polychrosis deltophora based on a single female from Mozambique, Magude. The type specimen which unfortunately lacks the abdomen, was photographed and figured by Razowski & Krüger (2007). The specimens that we assign to this species agree with Meyrick’s original description and with the photo of the type. They were located in the BMNH collection among South African unidentified Tortricidae , and had been mixed up with specimens of Paraeccopsis exhilarata .

Paraeccopsis insellata ( Meyrick, 1920) ( Figs. 18 , 42 , 51 )

Argyroploce insellata Meyrick, 1920: 66 .

Paraeccopsis insellata, Aarvik 2004 , figs. 73, 74.

Type material. Holotype ♀, labelled: TYPE; Afrique or. anglaise Pori: Mbuyuni 1100 m, iii.1912, Alluaud & Jeannel; Argyroploce insellata Meyr. , type; HOLOTYPE Argyroploce insellata Meyrick ♀, Genital prep. 99024 L. Aarvik ( MNHN).

Additional material examined. KENYA: Rift Valley, Lake Bogoria 0˚20’46’’N, 36˚ 04’E, 1♂, 21.xi.2006, D.J.L. Agassiz, genitalia slide L. Aarvik 2013.019; 1♀, same locality, 20.xi.2006, genitalia slide L. Aarvik 2013.020; 1♀, Rift Valley, L. Baringo, 1000 m, 0˚36’N, 36˚00’E, 17.viii.2007, genitalia slide L. Aarvik 2013.021 ( DJLA).

Diagnosis. Paraeccopsis insellata has the forewing ochreous brown with blackish brown markings, which makes it nearly identical externally to P. deltophora and also similar to the light form of P. pseudoinsellata sp. n. The male genitalia ( Fig. 42 ) has a short uncus and resembles the one found in P. deltophora ( Fig. 41 ), whereas the shape of the valva is basically the same as in P. pseudoinsellata sp. n. ( Fig. 43 ). However, in P. deltophora and P. pseudoinsellata sp. n. the valva has an extra medial excavation which is not present in P. insellata . In the female genitalia it differs from P. deltophora and P. pseudoinsellata by the narrower excavation of the posterior edge of sternite 7 and by the short sterigma.

Distribution. Aarvik (2004) recorded P. insellata from Botswana, Gambia, India, Kenya, Nigeria and Tanzania; Razowski (2008) recorded it from Mozambique. However, we now consider Paraeccopsis inflicta (Meyrick) and P. atricapsis (Meyrick) as valid species (see below). This implies that P. insellata has so far only been reliably recorded from Kenya.

Remarks. Aarvik (2004) synonymised Paraeccopsis inflicta ( Meyrick, 1920) and P. atricapsis (Meyrick, 1930) with P. insellata . The examination of a larger number of species has shown that specific differences in the genitalia of Paraeccopsis females in some cases are small, and that P. inflicta and P. atricapsis should be treated as valid species. Paraeccopsis inflicta was described from a single female from India, and P. atricapsis from a single female from Nigeria. The two species and their genitalia were figured by Clarke (1958, pl. 288). The female genitalia of P. insellata differ from those of both P. inflicta and P. atricapsis by narrower excavation of the posterior edge of sternite 7. Paraeccopsis inflicta ( Meyrick, 1920) stat. n. and P. atricapsis (Meyrick, 1930) stat. n. are herewith removed from synonymy under P. insellata . The recently collected series of P. insellata containing two females and a male, has shown that Aarvik (2004) confused the male of P. pseudoinsellata sp. n. (described below) with P. insellata . His figures of the female genitalia are correct as they show the genitalia of the holotype of P. insellata . The figures of the male genitalia represents P. pseudoinsellata sp. n. Until now only the type specimen of P. insellata was known. The single male has lost the tip of the phallus, probably eaten by ants in the trap.

Paraeccopsis pseudoinsellata Aarvik , sp. n. ( Figs. 19, 20 , 43 , 50 )

Type material. Holotype ♂, KENYA: Rift Valley Province: Laikipia, Mpala Research Station, 1710 m., 1♂, 26–28.xi.2008, L. Aarvik, D. Agassiz, A. Kingston, genitalia slide NHMO 2246 ( NHMO). Paratype (1). KENYA: Rift Valley, Turi , 8000 ft., 1♀, 9.xi.1999, D.J.L. Agassiz, genitalia slide L. Aarvik 2012.012 ( DJLA).

Additional specimen examined. TANZANIA: Morogoro Distr. & Town, 550–600 m, 1♂, 23.iii.1993, L. Aarvik, genitalia slide L. Aarvik 2525 ( NHMO).

Description. Adult. Male ( Fig. 19 ). Head: Beige; antenna light brown, short ciliate; labial palpus beige, external surface with some brown scales, 1.5 times diameter of eye. Thorax: Beige, with dark brown transverse bands; crest brownish red, cream in middle. Legs light beige, fore tarsi with grey rings. Wingspan 15.0 mm. Forewing ground colour cream; heavily suffused with blackish brown and dark grey, also some reddish brown; dark suffusion dominates median part of wing, forming costal blotch; white discal spot below it; dark suffusion present in apical area and also as an oblique band from mid-termen towards costa; tornal area cream. Cilia browngrey, cream at tornus. Hindwing grey, becoming lighter towards base, veins darker; cilia dirty white, with single dark cilia line. Abdomen: Genitalia ( Fig. 43 ). Uncus high, slender, triangular, gradually becoming narrower towards knob-like, spiny termination; socii shaped as narrow lobes; valva rather straight, with medial oval excavation in addition to the normal basal excavation; edge of medial excavation sclerotised, in particular in the right valva where the dorsal edge is widened to a plate carrying numerous long spines; left valva with a few spines at the distal edge of the medial excavation; cucullus about half the length of valva, apically rounded; phallus sinuous, no carinae or cornuti.

Female. Head and thorax: Similar to male. Abdomen: Genitalia ( Fig. 50 ). Posterior edge of sternite 7 with wide V-shaped excavation; sterigma shaped as an oblique tube, becoming narrower posteriorly; sclerotised portion of ductus bursae relatively long and slender, reaching beyond anterior edge of sternite 7; signum an irregular sclerite with 2 anteriorly directed projections of dissimilar size.

Variation. The male from Tanzania ( Fig. 20 ), not included in the type series, has much lighter forewing maculation than the two types. This specimen was also figured (as P. insellata ) by Aarvik (2004), and its male genitalia matches those of the holotype closely. The only difference noted is that the Tanzanian specimen has the tip of the valva slightly more pointed. We ascribe this to variation.

Diagnosis. The male genitalia resemble those of Paraeccopsis insellata , but the valva of P. pseudoinsellata sp. n. has a medial oval excavation in addition to the normal basal excavation ( Fig. 43 ). The medial excavation is lacking in P.insellata ( Fig. 42 ). In the female genitalia ( Fig. 50 ) the sterigma is an oblique tube. In P. insellata ( Fig. 51 ) the sterigma is much shorter. As few specimens are known, little is known about external variation of this species and P. t u r i sp. n. which occurs in the same area. It is recommended that genitalia slides are made for identification.

Distribution. Known from Rift Valley Province in Kenya and from eastern Tanzania. Material from Gambia listed by Aarvik (2004) under P. insellata belongs to neither P. insellata nor P. pseudoinsellata , sp. n. Etymology. The name reflects the similarity with Paraeccopsis insellata , especially in the male genitalia, and also that the two species were confused.

Paraeccopsis phoeniodes ( Meyrick, 1921) ( Figs. 21 , 44 , 52 )

Argyroploce phoeniodes Meyrick, 1921: 59 ; Razowski & Krüger 2007: 120, fig. 68. Paraeccopsis phoeniodes, Razowski 2008: 62 .

Material examined. 1♂, KENYA: Rift Valley Province: Turi , 9.ix.1999, D.J.L. Agassiz, genitalia slide L. Aarvik 2013.002 ( DJLA); 1♀, Rift Valley Province: Rongai, 6.i.2000, D.J.L. Agassiz, genitalia slide L. Aarvik 2013.016 ( DJLA); 1♀, Rift Valley Province: Gilgil, 2100 m, 0˚32’S, 36˚22’E, 26.xi.2005, D.J.L. Agassiz; 1♂, same locality, 3.ix.2006 ( DJLA).

Diagnosis. This species can be distinguished superficially from congeners by the more elongate wing shape and the reddish forewing colour ( Fig. 21 ). The forewing pattern elements are the same as in other Paraeccopsis species, but the fasciae are more narrow. The male genitalia ( Fig. 44 ) have a slender tegumen and weakly demarcated uncus; a slender valva, with a short rounded cucullus; and the field of spines on the left valva is larger than the one on the right valva. The female genitalia ( Fig. 52 ) have a large and round ostium, and a strongly sclerotised sterigma, narrowing posteriorly, divided into two laterally diverging ‘wings’.

Distribution. Mozambique, Kenya and Republic of South Africa.

Remarks. The holotype female from Mozambique was figured by Razowski & Krüger (2007). Unfortunately, it lacks the abdomen. Razowski (2008) mentioned one additional specimen from South Africa, Illwo Beak. Paraeccopsis phoeniodes is distinct externally, and based on this, we identify the Kenyan material as P. phoeniodes .

Figures

FIGURES 5 – 12. Adults of Paraeccopsis. 5 – 10. P. v ar i e g an a, sp. n., showing variation. 11. P. exhilarata (Meyrick). 12. P. windhoeca Razowski.

FIGURES 34 – 37. Male genitalia of Paraeccopsis. 34. P. variegana, sp. n. 35. P. exhilarata (Meyrick). 36. P. windhoeca Razowski. 37. P. tanzanica Aarvik, sp. n.

FIGURES 45 – 47. Female genitalia of Paraeccopsis. 45. P. variegana, sp. n. 46. P. exhilarata (Meyrick). 47. P. addis Aarvik, sp. n.

FIGURES 13 – 20. Adults of Paraeccopsis. 13. P. tanzanica Aarvik, sp. n. 14. P. addis Aarvik, sp. n. 15. P. turi Aarvik, sp. n. 16. P. botswanae Aarvik, sp. n. 17. P. deltophora (Meyrick). 18. P. insellata (Meyrick). 19. P. pseudoinsellata Aarvik, sp. n., dark female from Kenya. 20. P. pseudoinsellata Aarvik, sp. n., light male from Tanzania.

FIGURES 38 – 41. Male genitalia of Paraeccopsis. 38. P. addis Aarvik, sp. n. 39. P. tur i Aarvik, sp. n. 40. P. botswanae Aarvik, sp. n. 41. P. deltophora (Meyrick).

FIGURES 48 – 50. Female genitalia of Paraeccopsis. 48. P. t ur i Aarvik, sp. n. 49. P. deltophora (Meyrick). 50. P. pseudoinsellata Aarvik, sp. n.

FIGURES 42 – 44. Male genitalia of Paraeccopsis. 42. P. insellata (Meyrick). 43. P. pseudoinsellata Aarvik, sp. n. 44. P. phoeniodes (Meyrick).

FIGURES 51 – 54. Genitalia of Paraeccopsis and Age. 51. P. insellata (Meyrick). 52. P. phoeniodes (Meyrick). 53. Age onychistica male. 54. Age onychistica female.

FIGURES 21 – 25. Adults and genitalia of Paraeccopsis and Endotera. 21. Adult of Paraeccopsis phoeniodes (Meyrick). 22. Adult of Endotera cyaneana Agassiz, sp. n. 23. Freshly emerged specimen of Endotera cyaneana Agassiz, sp. n. on leaves of Acacia drepanolobium. 24. Male genitalia of Endotera cyaneana Agassiz, sp. n. 25. Female genitalia of Endotera cyaneana Agassiz, sp. n.