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Sakaija japonica ( Rathbun, 1932 )

Ng, Peter K. L. & Forges, Bertrand Richer De, 2015, Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific, Raffles Bulletin of Zoology 63, pp. 110-225 : 169

publication ID

 https://doi.org/10.5281/zenodo.5384590

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 lsid:zoobank.org:pub:40BCDD62-D35E-46D1-95A3-2CC0DF219DEE

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scientific name

 Sakaija japonica ( Rathbun, 1932 )
status

 

Sakaija japonica ( Rathbun, 1932) View in CoL

( Figs. 28A–C View Fig , 31A–C View Fig , 33A–G View Fig , 37K View Fig , 40J View Fig , 42F View Fig , 46G View Fig , 51A, B View Fig , 52J View Fig , 54J View Fig , 56H View Fig )

Maja sp. – T. Sakai, 1932: 50, pl. 2, fig. 5, text-fig. 5.

Maja japonica Rathbun, 1932: 33 View in CoL . – T. Sakai, 1965: 83, pl. 37, fig. 1. – Serène, 1968: 57. – Takeda & Miyake, 1969: 512, pl. 17, fig. A, B, pl. 18, fig. B. – Takeda, 1973: 42. – Takeda, 1975: 127. – T. Sakai, 1976: 238, pl. 82, fig. 1; text-figs 126a, b. – Miyake, 1983: 44. – Dai et al., 1986: 150, pl. 18(1), text-fig. 76. – Griffin & Tranter, 1986: 216. – Huang, 1989: 339. – Dai & Yang, 1991: 135, pl. 18(1), text-fig. 76. – Takeda, 1993: 43. – Huang, 1994: 582. – Muraoka, 1998: 27. – Ng et al., 2001: 12. – Takeda, 2001: 235. – Maramura & Kosaka, 2003: 34. – Yang et al., 2008: 780. – Ng et al., 2008: 117 (list).

Maja japonica Yokoya, 1933: 157 View in CoL , text-fig. 56.

Maja nipponensis T. Sakai, 1934: 297 View in CoL , text-fig. 11. – T. Sakai, 1936: 100, text-fig. 46. – T. Sakai, 1938: 299, pl. 38, fig. 1, text-fig. 41.

Maja sakaii View in CoL – Miyake, 1983: 44, pl. 15 fig. 5.

Material examined. Holotype: male (16.2 × 13.0 mm) ( USNM 48252 About USNM ), off Honshu, Japan, coll. RV Albatross, 7 May 1900. Others: Japan – 1 male (18.7 × 14.6 mm) ( NHM 1961.6.5.120), Sagami Bay, Japan, coll. T. Sakai. — 1 male (with bopyrid) (15.6 × 13.4 mm), 1 female (20.3 × 15.4 mm) (NSMT-Cr 13177), Tosa Bay , coll. RV Kotoka-Maru, 21 January 1998. — 1 male (with bopyrid) (NSMT-Cr 1132), off Tsushima Islands , station 10, 33°57.7’N 129°11.6’E, sand and shell substrate, coll. Tsushima Expedition, 26 July 1968. — 1 male, 1 female ( USNM 120722 About USNM ), off Ashizuri Peninsula , Shikoku Island , 300 m, coll. T. Sakai & K. Sakai, 27–31 March 1966. — 1 juvenile female (carapace only) ( USNM 48483 About USNM ), station 4893, 10– 20 miles southwest of Goto Island , Eastern Seas , 106 fathoms, coll. RV Albatross, 9 August 1906. — 1 male, 2 females ( SMF), Tosa Bay , Kochi, Usa, coll. RV Toyohata-Maru, K. Sakai. Taiwan – 1 male (22.3 × 17.8 mm) ( ZRC 2013.1267 View Materials ), Tashi fishing port, Ilan County, northern Taiwan , coll. 11 July 1989. — 1 broken male (20.8 × 16.7 mm) ( ZRC 2013.2371 View Materials ), Tashi fishing port, Ilan County , northern Taiwan , coll. S.-H. Wu, 1997. — 1 ovigerous female (20.4 × 16.4 mm) ( ZRC 2013.1270 View Materials ), Tashi fishing port, Ilan County , northern Taiwan , coll. S.-H. Wu, 23 January 1999. — 1 male (22.6 × 17.9 mm) ( ZRC 2013.1269 View Materials ), Tashi fishing port, Ilan County , northern Taiwan , coll. S.-H. Wu, 8 March 1999. — 1 male (18.5 × 13.9 mm) ( ZRC 2013.1268 View Materials ), Tashi fishing port, Ilan County , northern Taiwan , coll. Hwang, 18 July 2011. GoogleMaps

Diagnosis. Carapace branchial area strongly inflated, rounded; with relatively short lateral and branchial spines ( Figs. 28A–C View Fig , 31A, B View Fig ). Pseudorostral spines short, diverging ( Fig. 37K View Fig ). Supraorbital eave strongly curved; antorbital spine short, triangular, sharp; intercalated spine triangular, separated from adjacent spines by large gaps; postorbital spine longest, triangular, pointed anteriorly; hepatic spine short pointed outwards ( Figs. 28A–C View Fig , 37K View Fig ). Lateral margin with 2 short spines, posterior one larger (often becoming tubercles in larger specimens); branchial region with 1 low spine or tubercle; median row with 4 low spines or tubercles: 2 gastric (posterior one larger), 1 cardiac, 1 intestinal; 2 short spiniform granules on posterior carapace margin ( Figs. 28A–C View Fig , 31A, B View Fig ). Basal antennal article long with several granules, 2 strong distal spines ( Fig. 40J View Fig ). Ischium of third maxilliped weakly granulated ( Fig. 46G View Fig ). Ambulatory legs relatively short ( Figs. 31C View Fig , 56H View Fig ). G1 distinctly curved, distal part long, tapering with very long setae ( Fig. 33A–G View Fig ).

Remarks. Despite its age, the type specimen is in a relatively good condition although the G1 appears rather frail and not easy to study. However, it agrees well with those from Japan and Taiwan, and we are confident of their conspecificity.

For a detailed discussion of its taxonomy, see remarks for S. sakaii .

In Japan, S. japonica has been reported from depths of between 85 and 120 m. Outside Japan, it has been recorded from Hong Kong ( Griffin & Tranter, 1986), although their specimens should be re-examined .

Dai A-Y, Yang S-L, Song Y-Z & Chen G-X (1986) Crabs of the China Seas. China Ocean Press, Beijing. 11 + 642 pp.

Dai A-Y & Yang S-L (1991) Crabs of the China Seas. China Ocean Press, Beijing and Springer-Verlag, Berlin, 21 + 608 pp., figs. 1 - 295, pls. 1 - 74

Griffin DJG & Tranter HA (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie, 39 C 4: 1 - 335, 22 pls.

Huang L (1989) [Brachyura (Dromiacea, Gymnopleura, Oxystomata, Oxyrhyncha, Corystoidea)]. In: Dong Y (ed.) Fauna of Zhejiang. Zhejiang Science and Technology, Zhejiang. Pp. 280 - 348.

Huang Z-G (ed.) (1994) Brachyura. In: Marine species and their distributions in China's Seas. Department of Ocean Management and Monitoring State Oceanic Administration. China Ocean Press, Beijing. Pp. 576 - 600.

Miyake S (1983) Japanese Crustacean Decapods and Stomatopods in color. Vol. II, Brachyura (Crabs). Hoikusha Publishing Co Ltd., 277 pp.

Muraoka K (1998) Catalogue of the Brachyuran and Anomuran Crabs donated by Prof. Dr. Tune Sakai to the Kanagawa Prefectural Museum. Catalogue of the Collection in the Kanagawa Prefectural Museum of Natural History, Odawara, Kanagawa, Japan, No. 11: 1 - 67.

Ng PKL, Wang C-H, Ho P-H & Shih H-T (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum. Special Publication Series, Number 11: 1 - 86.

Ng PKL, Guinot D & Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286.

Rathbun MJ (1932) Preliminary descriptions of new species of Japanese crabs. Proceedings of the Biological Society of Washinginton, 45: 29 - 38.

Sakai T (1932) Notes on some rare materials of Japanese Oxyrhyncha. Science Reports of the Tokyo Bunrika Daigaku, Section B, 1 (4): 41 - 59, figs. 1 - 8, pls. 2, 3.

Sakai T (1934) Brachyura from the coast of Kyusyu, Japan. Science Reports of the Tokyo Bunrika Daigaku, Section B, 1 (25): 281 - 330, figs. 1 - 26, pls. 17, 18.

Sakai T (1936) Crabs of Japan. 66 plates in life colours with descriptions. Sanseido, Tokyo, pp. 1 - 239, 1 - 12 [literature cited], 1 - 27 [bibliography and index], pls. 1 - 66. [dated 1935 but published in 1936]

Sakai T (1938) Studies on the Crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo, Tokyo, pp. 193 - 364, figs. 1 - 55, pls. 20 - 41, table I.

Sakai T (1965) The Crabs of the Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen Co., Tokyo. i - xvi + 1 - 206 pp. [English section], figs. 1 - 27, pls. 1 - 100, 1 - 92 [in Japanese] + 1 - 26 [Bibliography and Index in English] + 27 - 32 [Index in Japanese] pp., 1 map.

Sakai T (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd., Tokyo. In three volumes; English Text, pp. xxix + 773 pp., Japanese Text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251.

Serene R (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. Special Publication of the Singapore National Academy of Science, No. 1: 33 - 120.

Takeda M & Miyake S (1969) Crabs from the East China Sea. III. Brachygnatha Oxyrhyncha. Journal of the Faculty of Agriculture, Kyushu University, 15 (4): 469 - 521, figs. 1 - 12, pls. 17, 18.

Takeda M (1973) Report on the crabs from the sea around the Tsushima Islands collected by the research vessel Genkai for the trustees of the National Science Museum, Tokyo. Bulletin of the Liberal Arts & Science Course, Nihon University School of Medicine, 1: 17 - 68.

Takeda M (1975) Brachyura. In: Utinomi H (ed.) Aquatic Invertebrates. Gakken Chukosei zukan, 9: 120 - 149 (fig. with Latin names), 188 - 331 (descriptions). First Edition. Gakken, Tokyo. [Second Edition in 1983 as The Aquatic Lower Animals of Japan without changes].

Takeda M (1993) Catalogue of crab specimens (Crustacea: Decapoda: Brachyura) (Part 1). National Science Museum, Tokyo, 81 pp.

Takeda M (2001) Annotated list of crabs from Tosa Bay, Southwest Japan, collected by the R / V Kotaka Maru during the years 1997 - 2000. In: Fujita T, Saito H & Takeda M (eds) Deep-sea fauna and pollutants in Tosa Bay. National Science Museum Monographs, 20: 217 - 262.

Yang S, Chen H & Jiang W (2008) Crustacea: Decapoda Brachyura. In: Liu R (ed.), Checklist of Marine Biota of China Seas. Institute of Oceanology, Chinese Academy of Sciences. Pp. 761 - 810.

Yokoya Y (1933) On the distribution of decapod crustaceans inhabiting the continental shelf around Japan, chiefly based upon the materials collected by the S. S. Soyo-Maru, during the year 1923 - 1930. Journal of the College of Agriculture, Imperial University of Tokyo, 12 (1): 1 - 226, figs. 1 - 71, tabs. 1 - 4.

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Fig. 28. General habitus, Sakaija species. A, S. japonica, holotype male (16.2 × 13.0 mm) (USNM 48252), Japan; B, S. japonica male (22.6 × 17.9 mm) (ZRC 2013.1269), Tashi fish port, Ilan County, northern Taiwan; C, S. japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; D, S. sakaii (as Maja sakaii, after T. Sakai 1976: pl. 85 fig. 2); E, S. sakaii (as Maja sakaii, after T. Sakai, 1976: text-fig. 125a); F, S. sakaii (as Maja sp., after T. Sakai, 1932: text-fig. 5); G, S. sakaii, male (9.0 × 6.6 mm) (NSMT-Cr 8094), Japan.

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Fig. 31. Frontal and lateral views of carapaces, and right first ambulatory legs, Sakaija species. A–C, S. japonica male (22.6 × 17.9 mm) (ZRC 2013.1269), Taiwan; D–F, S. serenei n. sp., paratype male (17.7 × 14.6 mm) (ZRC 2013.1262), Philippines.

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Fig. 33. Left G1s, Sakaija species. A–C, S. japonica, holotype male (16.2 × 13.0 mm) (USNM 48252), Japan; D–F, S. japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; G, S. japonica (as Maja japonica, after T. Sakai, 1976: text-fig. 126a); H, S. sakaii (as Maja sakaii, after Sakai, 1976: text-fig. 125b); I, J, S. sakaii, male (9.0 × 6.6 mm) (NSMT-Cr 8094), Japan; K–M, S. serenei n. sp., paratype male (14.3 × 10.9 mm) (ZRC 2013.1255), Philippines; N, O, S. africana (as Maja africana, after Griffin & Tranter, 1986: fig. 73c, d); P–R, S. africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; S–U, S. santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu. Scale bars = 1.0 mm [D, K, P–R]; 0.5 mm [A, E, F, I, J, L, M, S–U]; 0.25 mm [B, C].

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Fig. 37. Frontal regions of carapaces. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; J, Holthuija poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea; K, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; L, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; M, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; N, Sakaija longispinosa n. sp., holotype ovigerous female (11.4 × 8.6 mm) (NMV J63792), Australia; O, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu.

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Fig. 40. Antennae, antennules and epistomes. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; L, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; M, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; N, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; O, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan.

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Fig. 42. Epistomes. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; C, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; E, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; F, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; G, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; H, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan.

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Fig. 46. Third maxillipeds. A, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; B, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; C, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; D, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; E, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; F, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; G, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; H, Sakaija sakaii, male (9.0 × 6.6 mm) (NSMT-Cr 8094), Japan; I, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; J, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; K, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; L, Sakaija longispinosa n. sp., paratype ovigerous female (14.2 × 10.7 mm) (NMV J63197), Australia; M, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; N, Planaja plana n. gen. n. sp., paratype male (38.3 × 30.7 mm) (ZRC 2013.1370), Philippines; O, Ovimaja compressipes, holotype female (51.2 × 40.5 mm) (NHM 1860.15), China; P, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan.

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Fig. 51. Anterior thoracic sternums and abdomens. A, Sakaija japonica, holotype male (16.2 × 13.0 mm) (USNM 48252), Japan; B, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; C, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; D, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; E, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; F, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; G, Planaja plana n. gen. n. sp., paratype male (38.3 × 30.7 mm) (ZRC 2013.1370), Philippines; H, Ovimaja compressipes, holotype female (51.2 × 40.5 mm) (NHM 1860.15), China; I, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan.

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Fig. 52. Male sterno-abdominal cavities. A, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; B, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; C, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; D, Paramaja gibba n. sp., male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; F, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; G, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; H, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; I, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; L, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan.

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Fig. 54. Chelae. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., paratype male (76.5 × 61.3 mm) (ZRC 2001.0577), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; G, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; H, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; L, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; M, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; N, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; O, Ovimaja compressipes, holotype female (51.2 × 40.5 mm) (NHM 1860.15), China; P, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan.

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Fig. 56. First ambulatory dactyli and propodi. A, Paramaya spinigera, dried female (70.1 × 58.8 mm) (KPM NH4195), Japan; B, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; C, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; E, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan [figure laterally inverted]; F, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; G, Holthuija poorei, n. sp. paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea; H, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; I, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines [figure laterally inverted]; J, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; K, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; L, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan.

RV

Collection of Leptospira Strains
NHM

University of Nottingham
T

Tavera, Department of Geology and Geophysics
SMF

Forschungsinstitut und Natur-Museum Senckenberg

Kingdom

Animalia

Phylum

Arthropoda

Class

Malacostraca

Order

Decapoda

Family

Majidae

Genus

Sakaija

Specimens