Achelia Hodge, 1884

Lucena, Rudá Amorim & Christoffersen, Martin Lindsey, 2018, An annotated checklist of Brazilian sea spiders (Arthropoda: Pycnogonida), Zootaxa 4370 (2), pp. 101-122 : 104

Achelia Hodge, 1884

Achelia besnardi Sawaya, 1951: 271 –279, figs 1–2 ( Brazil: Rio de Janeiro, 39–46 m) .— Stock, 1992: 116 ( Brazil: off Espírito Santo, 59 m) .

Syn.: Achelia sawayai f. besnardi — Stock, 1986.

Holotype: Not designated.

Type locality: São Tomé shoal , Rio de Janeiro, Brazil .

Ecological note: Found among hydroids and algae.

Achelia columnaris Stock, 1992: 116 –118, 139, figs 9–18 (Brazil: Rio de Janeiro, 34 m). Holotype: ZMA.PYC.P.1023.

Type locality: Rio de Janeiro (22°22'S 41º32'W), Brazil.

Achelia gracilis Verrill, 1900 — Stock, 1992: 118 (Brazil: Espírito Santo, 28 m). Syn.: Achelia (?) gracilis —Verrill, 1900.

Ammothea gracilis —Cole, 1904.

Ammothea (Achelia) gracilis —Giltay, 1934.

Achelia (Pigrolavatus) gracilis —Fry & Hedgpeth, 1969. Holotype: Unknown.

Type locality: Flatts Inlet , Bermuda.

Achelia sawayai Marcus, 1940 c: 81 –86, figs 10a–f, 17 (Brazil: São Paulo, shallow water).— Stock, 1992: 118 (Brazil: São Paulo, off Rio de Janeiro, Espírito Santo, intertidal to 98 m).— Varoli, 1996: 42 (Brazil: São Paulo).— Morgado & Tanaka, 2001: 177 [tab.] (Brazil: São Paulo, 0.6–5 m).— Vieira et al., 2012: 164 (Brazil: Saint Peter and Saint Paul Archipelago).

Syn.: Achelia (Pigrolavatus) sawayai —Fry & Hedgpeth, 1969.

Achelia sawayai f. typica — Stock, 1986.

Non Achelia sawayai f. besnerdi — Stock, 1986 (= A. besnardi Sawaya, 1951 ).

Holotype: Not designated.

Type locality: Baía de Santos , São Paulo, Brazil .

Ecological notes: On seaweeds, rocky shore. On large boulders, amongst ascidians, bryozoans and sponges. On

Dictyota sp., Sargassum cymosum C. Agardh , and Sargassum rigidulum Kuetzing. On bryozoan Schizoporella

errata (Walters) .

Figures

FIGURE 1. Map of Brazil, in light grey. Remaining countries in South America are in dark grey. Abbreviations, States in Northern Region: AC, Acre; AM, Amazonas; AP, Amapá; PA, Pará; RO, Roraima; RR, Rondônia; TO, Tocantins. States in Northeastern Region: AL, Alagoas; BA, Bahia; CE, Ceará; MA, Maranhão; PB, Paraíba; PE, Pernambuco; PI, Piauí; RN, Rio Grande do Norte; SE, Sergipe. States in Southeastern Region: ES, Espírito Santo; MG, Minas Gerais; RJ, Rio de Janeiro; SP, São Paulo.States in Central-West Region:DF, Distrito Federal; GO, Goiás; MS, Mato Grosso do Sul; MT, Mato Grosso. States in Southern Region: PR, Paraná; RS, Rio Grande do Sul; SC, Santa Catarina.Rivers:AmR, Amazon River; DoR, Doce River; PaR, Parnaíba River; SaR, São Francisco River.

FIGURE 2. Map of Brazilian littoral showing sampling sites for pycnogonids according to studies published. Due to close location of sites sampled in the same study not all sites are shown.Studies are refered by the first author's name.(A) studies by local researchers; (B) studies by visiting researchers.