Cochlostoma (T.) gracile gracillimum ( Wagner, 1901 )

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 28-29

Cochlostoma (T.) gracile gracillimum ( Wagner, 1901)

Figs 15H , 21 (green dots), 23

Pomatias gracilis var. gracillima Wagner, 1901: 64 .

Syntypes

BOSNIA AND HERZEGOVINA • 1 ♀, 1 ♂; “ Bočac zwisch. Jajce und Banjaluka ”; 1 Aug. 1899; “ Auritus gracile gracillimus, Wagner det.”; Sturany leg.; NHMW3230 • 4 ♀♀; “ Bočac zw. Jajce und Banjaluka ”; as Pomatias gracilis var. gracillima ; MIZPASW6866 .

Other specimens

BOSNIA AND HERZEGOVINA • 2- Vrbac valley ; 44.6833° N, 17.1783° E; 2013; J. Grego leg.; coll. Grego (Horna Micina Slovakia) .

Type locality

BOSNIA AND HERZEGOVINA • 1- Republika Srpska, Bočac.

Description

SHELL. Closely spaced riblets on last part of protoconch. Teleoconch spotless, with strong to moderately strong and rounded ribs. Rather weak lip with columellar lobe quite abruptly bent inwardly to cover umbilicus.

MEASUREMENTS. 2 ♀♀: whorls= 7.8–8.6, H = 6.8–7.5 mm, H/W= 2.81–2.94, roundness =0.2–0.21, ribs incl.= 58–61°, apert. incl.= 17–20°, ribs/mm 1 st wh.= 10–10, ribs/mm 4 th wh.= 10–10.

FEMALE GENITAL ORGANS. As in C. (T.) stussineri .

Remarks

We provisionally keep it as a subspecies of “ gracile ” because of insufficient data to revise its status in more detail. The name “ gracillimum ” is a declination in Latin of the superlative of “ gracilis ” and can be translated as ‘really slender’ but the analysed specimens do have a H/W ratio similar to that of the nominotype. In Wagner’s collection, there are specimens from the type locality and from Livno, another locality in Bosnia, situated more than 100 km south (MIZPAS-W6867), which might belong to the same taxon. The p-distance of this taxon with the nominotyical sample of the species is 1.4%.

Figures

Fig. 15. Shell and female genitals of clade A samples with amplified 16S which share the same female genital set-up. A. NFS158, Tisovec, SLO (WdM-6748). B. Cochlostoma (T.) reitteri (Boettger, 1880), Kapela Pass, HR (WdM-6882). C. C. (T.) stussineri (Wagner, 1897), Rakow Skocjan, SLO (WdM-6918). D. NFS064, Terminillo V. Dell’Inferno, I (EZ-1067). E. C. (T.) patulum (Draparnaud, 1801), Cirque D’Archiane, FR (RMNH-117454). F. NFS153, 3 km SE of Oltari, HR (HNHM-100381). G. C. (T.) gracile gracile (Pfeiffer, 1849), Omiš 1, HR (WdM-6667). H. C. (T.) gracile gracillimum (Wagner, 1901), Vrbac Valley, BeH (Gr001).

Fig. 21. Distribution of the samples of Cochlostoma (T.) gracile (Pfeiffer, 1849): red dots= C. (T.) gracile gracile; green dots =C. (T.) gracile gracillimum (Wagner, 1901); cyan dots = C. (T.) gracile croaticum (Pfeiffer, 1870). A black center indicates samples with amplified 16S.