Cochlostoma (Turritus) stussineri ( Wagner, 1897 )

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 18-22

Cochlostoma (Turritus) stussineri ( Wagner, 1897)

Figs 14 (cyan dots), 15C, 16–17

Pomatias gracile var. stussineri Wagner, 1897: 46 , pl. 8 fig. 76a–b.

Lectotype (here designated)

SLOVENIA • ♀; 1- Nanos ober Präwald ; NHMWE 1262.

Other specimens

ITALY • 14- Mt Sabotino ; 45.9849° N, 13.6282° E; 2009; De Mattia leg.; WdM6954 .

SLOVENIA • 1- Mt Nanos , top, loc. Plesa (topotypical); 45.7981° N, 14.067° E; 2005; De Mattia leg.; WdM6727 • 1- Mt Nanos , 3.7 km NNW of Abram (topotypical); 45.8476° N, 14.0103° E; 2009; De Mattia leg.; WdM-6730 • 4- Divje Jezero ; 45.9827° N, 14.0277° E; 2009; De Mattia leg.; WdM6840 • 5- Solkan to Plava ; 46.0157° N, 13.6112° E; 2009; De Mattia leg.; WdM6695 • 6- Podlešče-Bainsizza ; 46.0574° N, 13.7271° E; 2009; De Mattia leg.; WdM6813 • 7- 2.4 NW of Kalce; 45.9074° N, 14.1647° E; 2009; De Mattia leg.; WdM6725 • 8- Planina ; 45.8297° N, 14.2506° E; 2000; Zallot leg.; EZ0125 • 9- 3 km S of Planina; 45.8300° N, 14.2606° E; 2000; Zallot leg.; EZ0126 • 10- near Logatec ; 45.8808° N, 14.2745° E; 2009; Zallot leg.; EZ0127 • 11- Vrhnika ; 45.9706° N, 14.3119° E; 2000; Zallot leg.; EZ0128 • 12- Rakow Skocjan ; 45.7947° N, 14.2895° E; 2009; Zallot leg.; WdM6918 • 13- Laška kukava ; 45.8907° N, 14.2889° E; 2008; De Mattia leg.; WdM6924 .

Type locality

SLOVENIA • Mount Nanos, Inner Carniola.

Description

SHELL. Spaced riblets on last part of protoconch. Teleoconch spotless, ribbed with ribs not very prominent. Moderately strong lip, with columellar lobe inwardly curved to cover the umbilicus.

MEASUREMENTS. 24 ♀♀: whorls=7.1–8.3, H= 6.4–9 mm, H/W =2.43–2.79, roundness =0.11–0.16, ribs incl.= 57–67°, apert. incl.= 9–24°, ribs/mm 1 st wh. =6–20, ribs/mm 4 th wh.=6–16.

FEMALE GENITAL ORGANS. Ventral connection of pedunculus of bursa copulatrix, short, oval-shaped seminal receptacle confined to ventral side of body, two to three loops close to apex of seminal receptacle and close connection of junction of uterus gland to connection between pedunculus of bursa copulatrix and distal oviduct.

Remarks

Wagner (1897: 46 [610]) firstly mentioned as locality “Nanos”. He also referred to specimens from “Krain. Idrizathal” as unconditionally belonging to this species. By the designation of NHMW E1262 as lectotype, Mount Nanos becomes the type locality. The specimens MIZPASW6869 (5 ♀♀), 4- Krain, Iška Schlucht b. Krimberg, SMF160611 (1 ♀), 4- Krain, Iška Schlucht b. Krimberg and SMF160613 (1 ♂) were also mentioned, but considered transitional to “ croatica ”, and cannot be considered syntypes. De Mattia et al. (2011) reported C. (T.) gracile from Mount Sabotino, Italy. In the light of the results of our analysis, this has to be corrected to C. (T.) stussineri .

The great variability of the number of ribs per mm and the variable strength and shape of the lip may indicate that not all populations assigned to this species are conspecific.

Figures

Fig. 14. Distribution of samples part of clade A: cyan dots= Cochlostoma (T.) stussineri (Wagner, 1897); red dots= C. (T.) patulum (Draparnaud, 1801); white dot =C. (T.) zawinkanum (Wagner, 1906). Black center for the samples with amplified 16S.