Cochlostoma (Turritus) patulum ( Draparnaud, 1801 )

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 22-26

Cochlostoma (Turritus) patulum ( Draparnaud, 1801)

Figs 14 (red dots), 15E, 19–20

Cyclostoma patulum Draparnaud, 1801: 39 .

Lectotype (here designated)

FRANCE • probably ♀; Hérault, Montpellier; “ Cyclostoma patulum Coll. Draparnaud 1820 XXVI /13” and “ Cochlostoma patulum (Drap.) Typus von Draparnaud!”; NHMW 14710/1.

Other specimens

FRANCE • 2- Cirque d’Archiane ; 44.7590° N, 5.4186° E; 1975; RMNH leg.; RMNH117454 • 3- Cassis ; 43.2157° N, 5.5389° E; 2009; Pocaterra leg.; EZ0457 • 4- Cirque de Navacelles ; 43.8945° N, 3.5115° E; 1978; de Winter leg.; A. de Winter-0882 • 5- Gorges de Galamus ; 42.8402° N, 2.4794° E; 1987; de Winter leg.; A. de Winter5601 .

Type locality

FRANCE • Hérault, Montpellier (“H.a.F.M.” in the original description =France meridionale. Wagner (1897) wrote: “Ich beurtheile diese Art nach Originalexemplaren Draparnaud’s von Montpellier in Südfrankreich, welche sich im k. k. naturhistorischen Hofmuseum in Wien befinden”.)

Description

SHELL. Closely spaced riblets on last part of protoconch. Teleoconch spotless, with strong ribs, slightly weaker and irregularly spaced approaching aperture. Strong and developed lip with columellar lobe abruptly inwardly curved to cover umbilicus.

MEASUREMENTS. 4 ♀♀: whorls= 7.2–8.1, H = 7.4–8.5 mm, H/W= 2.41–2.7, roundness= 0.12–0.13, ribs incl.= 61–64°, apert. incl.= 12–15°, ribs/mm 1 st wh.= 10–15, ribs/mm 4 th wh.= 11–13.

FEMALE GENITAL ORGANS. As in C. (T.) stussineri .

Remarks

Cochlostoma (T.) patulum from Cirque D’Archiane and from Luberon (GENBANK, GQ370416) do not cluster in the 16S analysis as do conspecific samples belonging to other species. The p-distance between them is 1.5%, higher than the one (1.2–1.3%) found between patulum and stussineri . For this reason, it seems appropriate to better define the type locality, here restricted to Montpellier ( France). Maybe ‘ancient’ DNA could be amplified from the lectotype, or a specimen of the same sample, to stabilize the taxonomy of the French Turritus .

Figures

Fig. 14. Distribution of samples part of clade A: cyan dots= Cochlostoma (T.) stussineri (Wagner, 1897); red dots= C. (T.) patulum (Draparnaud, 1801); white dot =C. (T.) zawinkanum (Wagner, 1906). Black center for the samples with amplified 16S.