Cochlostoma (T.) agriotes (Westerlund, 1879)

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 110-115

Cochlostoma (T.) agriotes (Westerlund in Paulucci, 1879)

Figs 97 (yellow dots), 98G, 100–101

Pomatias (patulus Drap. Subsp.) agriotes Westerlund in Paulucci, 1879: 20.

Types

Not seen.

Other specimens

ITALY • 1- Mt Cucco (10 km south along the same mountain range, here assumed as topotypical); 43.3646° N, 12.731° E; 2010; Zallot leg.; EZ0001 • 2- Gualdo Tadino ; 43.2302° N, 12.7957° E; 2013; Margelli leg.; EZ1139 • 9- Gole dell’Infernaccio ; 42.9397° N, 13.3159° E; 2013; Margelli leg.; EZ1142 .

Type locality

ITALY • Marche, Pesaro and Urbino, Fonte Avellana . Westerlund reported as localities “Italia at Suellana and Saorgio”. Paulucci already noticed that “Suellana” is the misspelled name of the hermitage Fonte Avellana. “Saorgio”, which is the second locality cited by Westerlund was once in Italy, but it is now a French village in the Maritime Alps which is inhabited by another taxon (locus typicus of C. (T.) simrothi ) .

Description

SHELL. Closely spaced riblets on last part of protoconch. Teleoconch spotless with more or less strong, often whitish, irregularly spaced ribs. Whitish powder on whorls. Rather weak lip with gently inwardly bent columellar lobe covering almost completely umbilicus.

MEASUREMENTS. 29 ♀♀: whorls= 7.1–9.2, H = 7.5–10.2 mm, H/W=2.54–3.04, roundness=0.1–0.18, ribs incl.= 52–64°, apert. incl. =15–24°, ribs/mm 1 st wh. =2–10, ribs/mm 4 th wh.=4–10.

FEMALE GENITAL ORGANS. As in C. (T.) affine .

Remarks

Cochlostoma (T.) agriotes is considered a subspecies of C. (T.) crosseanum in MolluscaBase (2023) but actually they have the same shell and female genital morphology. However, the Sicilian C. (T.) affine is equally similar and therefore, on morphological characters, it should be synonymized too. The p-distances between the samples of agriotes and crosseanum are very high (2.4–3.1%), higher than the one between the Sicilian affine and agriotes (1.7%). The assignment of populations to this species rather than to the nearby living C. (T.) crosseanum is purely tentative and based on geographical consideration.

Figures

Fig. 97. Distribution of samples in Clade B: green =Cochlostoma (T.) crosseanum (Paulucci, 1879); yellow= C. (T.) agriotes (Westerlund, 1879); red= C. (T.) affine (Benoit, 1876). Black center for samples with amplified 16S.