Cochlostoma (T.) nanum ( Westerlund, 1879 )

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 83-87

Cochlostoma (T.) nanum ( Westerlund, 1879)

Figs 72 (red dots), 73A–C, 74–76

Pomatias nanus Westerlund, 1879: 167-16 .

Pomatias clessini Hirc, 1881: 521–522 .

Types

Not seen.

Other specimens

AUSTRIA: 5- Tepke ; 46.3888° N, 14.5738° E; 2015; AL leg.; NHMW109000 /AL/00703.

SLOVENIA • 1- Mt Risnjak , 1400 m (topotypical); 45.4251° N, 14.6162° E; 2006; De Mattia leg.; WdM5666 • 2- Mt Snežnik 1764; 45.5884° N, 14.4478° E; 2010; De Mattia leg.; WdM7005 • 3- Mt Snežnik 1451; 45.5825° N, 14.4312° E; 2010; De Mattia leg.; EZ0960 • 4- Logarska Dolina ; 46.4274° N, 14.6572° E; 2009; Slapnik leg.; HNHM97239 .

Type locality

A generic “Croatia” in the original description. We define as type locality restricted: CROATIA • Mount Risnjak (see remarks) .

Description

SHELL. Closely spaced riblets on last part of protoconch. Teleoconch spotless, with weak or very weak ribs almost disappearing on last whorl and approaching aperture. Strong lip with columellar lobe either barely or abruptly inwardly bent, only partially covering umbilicus.

MEASUREMENTS. 13 ♀♀: whorls =7–8, H = 7–8.4 mm, H/W=2.4–2.64, roundness =0.1–0.15, ribs incl.= 53–61°, apert. incl.= 16–24°, ribs/mm 1 st wh.= 8–11, ribs/mm 4 th wh.=8–17.

FEMALE GENITAL ORGANS. Posterior connection of pedunculus of bursa copulatrix and junction of uterus gland moved far from connection between distal oviduct and pedunculus of bursa copulatrix. Seminal receptacle normally club-shaped, situated on dorsal side of body and with long and undefined basis/ distal oviduct.

Remarks

The type locality in the original description is rather undefined. We consider Mount Risnjak in the northern part of Croatia as type locality restricted by A.J. Wagner (1897: 54 [618]): “Die mir vom Autor übergebenen Originalexemplare stammen ohne nähere Fundortsangabe aus Croatien (ein bestimmter Fundort wird auch sonst vom Autor nirgends angeführt); dieselben erwiesen sich aber als vollkommen identisch mit den mir ebenfalls vorliegenden Originalexemplaren des P. clessini Hirc vom Berge Risnjak in Croatien und dürften wohl auch daher stammen [The original specimens were given to me by the author labeled ‘Croatia’ without any further information on where they were found (a specific site is also not mentioned anywhere else by the author); but it turned out that they are completely identical to the original specimens of P. clessini Hirc from the Risnjak mountain in Croatia that I also have and it may well be that they are from there]”.

For what is known, the species lives in a few scattered alpine localities not connected by suitable habitats. In the phylogenetic trees the nanum samples do not cluster together and the intraspecific p-distances are of the same magnitude of the ones with samples of the other species of the subclade.

Figures

Fig. 72. Distribution of samples of clade B: red =Cochlostoma (T.) nanum (Westerlund, 1879); cyan =C. (T.) elegans. Black center for the samples with amplified 16S.