Ceratocanthinae

Ballerio, Alberto, Gill, Bruce D. & Grebennikov, Vasily V., 2011, Illustrated overview and identification key to Cameroonian Ceratocanthinae beetles (Coleoptera: Scarabaeoidea: Hybosoridae) with description of four new species, Zootaxa 2892, pp. 1-24 : 22

Key to the described species of Cameroonian Ceratocanthinae

1 Protibiae broadly arcuate, outer margins at most finely serrate with weak tooth apically, no distinct teeth visible.......... 2

- Protibiae straight, with one or more distinct teeth on outer margin............................................... 9

2 Head with strong, large, dorsal trichome................................... Callophilharmostes fleutiauxi (Paulian)

- Head without strong, large, dorsal trichome, rarely with short setae............................................... 3

3 Genal canthus indistinct, no dorsal ocular area; small, flightless species (genus Baloghianestes )...................... 4

- Genal canthus distinct with dorsal ocular area; winged species.................................................. 7

4 Pseudoepipleura delimited by sharp carina, which is sometimes interrupted at middle; with pseudoepipleura abruptly folded inwards............................................................................................ 5

- Pseudoepipleura not delimited by sharp carina or, at most, by a hint of a sharp carina present near humeral area; with pseu- doepipleura almost indistinct............................................................................ 6

5 Elytral humeral area with a distinct, tubercle-like process ( Fig. 4 )................. Baloghianestes oribatidiformis n. sp.

- Elytral humeral area without tubercle or tubercle-like process ( Fig. 6 ).................. Baloghianestes korupensis n. sp.

6 Head with vertex having irregular raised longitudinal process. Setae clavate ( Fig. 5 )....... Baloghianestes lissoubai Paulian

- Head without raised, longitudinal process. Setae simple ( Fig. 3 ).......................... Baloghianestes anceps n. sp.

7 Dorsal surface of elytra and pronotum with several distinct tubercles ( Fig. 8 )........ Carinophilharmostes vadoni (Paulian)

- Dorsal surface of elytra and pronotum lacking carinae or tubercles.............................................. 8

8 Dorsal surface covered by short, thick setae ( Fig. 18 )............................ Petrovitzostes guineensis (Petrovitz)

- Dorsal surface lacking setation (low magnification: 20X)............................ Philharmostes badius (Petrovitz)

9 Length greater than 8 mm; genal canthus not reaching occipital area ( Figs. 9–11 )......... Congomostes hintelmanni n. sp.

- Length less than 8 mm; genal canthus reaching occipital area................................................. 10

10 Pseudoepipleura delimited at least partly by cariniform process................................................ 12

- Pseudoepipleura not delimited by cariniform process........................................................ 11

11 Elytra with irregular rows of small, shallow, comma-shaped punctures arranged transversely, basally and longitudinally along median and distal third ( Fig. 16 )....................................... Pseudopterorthochaetes endroedyi (Paulian)

- Elytra with large, horseshoe-shaped, shallow, transverse punctures ( Fig. 15 )......... Melanophilharmostes demirei Paulian

12 Surface of elytra and pronotum with fine, dense, simple punctures between fine, horseshoe-shaped punctures ( Fig. 17 )............................................................................ Melanophilharmostes zicsii (Paulian)

- Surface of elytra and pronotum smooth between large, shallow, ocellate punctures.. Melanophilharmostes burgeoni (Paulian)

Figures

FIGURE 3. Baloghianestes anceps n. sp., habitus. A—distended specimen, dorsal, B – D—rolled up specimen, ventral (B), dorsal (C), lateral (D).

FIGURE 4. Baloghianestes oribatidiformis n. sp., habitus. A—distended specimen, dorsal, B – D—rolled up specimen, ventral (B), dorsal (C), lateral (D).

FIGURE 5. Baloghianestes lissoubai Paulian, habitus. A—distended specimen, dorsal, B – D—rolled up specimen, dorsal (B), ventral (C), lateral (D).

FIGURE 6. Baloghianestes korupensis n. sp., habitus. A—distended specimen, dorsal, B – D—rolled up specimen, ventral (B), dorsal (C), lateral (D).

FIGURE 8. Carinophilharmostes vadoni (Paulian), habitus. A—distended specimen, dorsal, B – D—rolled up specimen, ventral (B), dorsal (C), lateral (D).

FIGURE 9. Congomostes hintelmanni n. sp., habitus. A—distended specimen, dorsal, B – D—rolled up specimen, ventral (B), dorsal (C), lateral (D).

FIGURE 10. Congomostes hintelmanni n. sp., A, C—parameres, dorsal view (A) and lateral (C), B—genital segment. Scale bar: 0.4 mm (A, C), 0.7 mm (B).

FIGURE 11. Congomostes hintelmanni n. sp., mouthparts. A – D—mandibles, dorsal (A, B) and ventral (C, D), E—labium, F—maxilla, G—details of galea brush, H—epipharynx.

FIGURE 15. Melanophilharmostes demirei Paulian, habitus. A—distended specimen, dorsal, B – D—rolled up specimen, ventral (B), dorsal (C), lateral (D).

FIGURE 16. Pseudopterorthochaetes endroedyi (Paulian), habitus. A—distended specimen, dorsal, B – D—rolled up specimen, ventral (B), dorsal (C), lateral (D).

FIGURE 17. Melanophilharmostes zicsii (Paulian), habitus. A—distended specimen, dorsal, B – D—rolled up specimen, ventral (B), dorsal (C), lateral (D).

FIGURE 18. Petrovitzostes guineensis (Petrovitz), distended specimen, dorsal. FIGURE 19. Pseudopterorthochaetes sp. A, distended specimen, dorsal.