Craspedochaeta transversa Czerny, 1903

Lonsdale, Owen & Marshall, Stephen A., 2006, Revision of the New World species of Craspedochaeta (Diptera: Clusiidae), Zootaxa 1291 (1), pp. 1-101 : 39-40

Craspedochaeta transversa Czerny, 1903

( Figs. 19 , 26 , 53, 54 , 61 , 74 , Map 6)

Craspedochaeta transversa Czerny, 1903: 103 . Type species of Craspedochaeta .

Czernyola transversa, Melander & Argo, 1924: 27 . Hennig, 1938: 131.

Redescription ( Fig. 19 , 26 )

Body length 2.8–3.3 mm. Bristles yellow (including bristles on basal section of costa). Small bristle occasionally present in front of anterior dorsocentral. Acrostichal bristle small. Scutum yellow with brown cruciate pattern (holotype and most specimens with transverse stripe incomplete). Scutellum and laterotergites brown. Pleuron and legs light yellow. Head predominantly yellow; anterior margin of frons and antenna light yellow; face white; back of head with one pair of stripes radiating from foramen; ocellar spot brown; face with one pair of faded lateral stripes; gena and parafacial white and silvery tomentose. Abdomen mostly black; tergite 1, usually tergite 2, and sometimes central spot on anterior margin of tergite 3 yellow; cerci and surstylus white. Wing clouded on distal half along R 2+3 and R 4+5 (darkest along R 2+3); R 4+5 and M 1 slightly convergent distally; distance between crossveins no more than length of posterior cross­vein.

Female

As described for male except as follows: “arms” of dorsal cruciate pattern not connected to central stripe; lateral stripes on face absent; epiproct yellow.

Male terminalia ( Figs. 53, 54 , 61 )

As described for C. pleuralis except as follows: cerci subequal in length to epandrium; surstylus straight with tip strongly tapered and with numerous minute bristles; anterior protuberance on pregonite highly reduced.

Female genitalia ( Fig. 74 )

Length of spermatheca 4.5 times width; distal segment hemispherical with apical depression; next most basal segment wide and ring­like; basal segment poorly sclerotized and cylindrical. Spermathecal duct thin and membranous; as long as basal segment of spermatheca. Ventral receptacle poorly sclerotized and spherical. Subterminal flagellum 12 times length of ventral receptacle.

Distribution: Argentina, Bolivia, Brazil, Costa Rica, Paraguay, Peru (Map 6).

Cotypes: PERU. “Peru” (2♀♀, NHMW) .

Additional material examined: ARGENTINA. Jujuy: Calilegua N.P., Estaca Bicaro, 900m, forest, 18–28.xii.1987, S.&J. Peck, Malaise trap (1♀, DEBU) . BOLIVIA. Depto La Paz, 8km S Chulumani, Apa Apa , ex. Malaise trap, 1950–2100m, 16°22’S, 67°30’W, 24.iii.2001, S.D. Gaimari (1♀, USNM) . BRAZIL. Terezópolis, Rio de Janiero , Servico Febre Amarela, M.E.S. Bras. (2♂♂ 4♀♀, USNM) , Roraima: Rio Uraricoera, Ilha de Maraca , 18–28.viii.1987, J.A. Rafael, L.S. Aquino, J. Vidal & E. Binda, Arm. Shannon (1♀, INPA) . COSTA RICA. La Suiza, P. Schild (1♀, USNM) , vii.1926 (1♀, USNM) , viii.1926 (1♀, USNM) , Guanacaste: 3km SE R. Naranjo, F.D. Parker, v.1992 (1♀, EMUS) , 22–24.vii.1992 (1♂, EMUS) .

Comments: Craspedochaeta transversa differs from similar species by having veins R 4+5 and M 1 convergent distally.

Figures

FIGURES 10–23: Head and thorax, dorsal colouration of the New World Craspedochaeta without an entirely dark notum; 10: C. xanthonotum spec. nov.; 11: C. festiva (Sóos, 1962); 12: C. loreto spec. nov.; 13: C. trivittata (Sóos, 1962); 14: C. weemsi spec. nov.; 15: C. pacaraima spec. nov.; 16: C. candida spec. nov.; 17: C. apsilutea spec. nov.; 18: C. ornata (Hennig, 1938); 19: C. transversa Czerny, 1903; 20: C. synneura (Hennig, 1938); 21: C. pleuralis (Williston, 1896); 22: C. pollostos spec. nov; 23: C. chela spec. nov, male.

FIGURES 24–26: Left wing, ventral (arrows indicate diagnostic characters mentioned in the key); 24: Craspedochaeta pleuralis (Williston, 1896); 25: C. synneura (Hennig, 1938); 26: C. transversa Czerny, 1903.

FIGURES 53–58: Male terminalia, external: Figs. 53–54: Craspedochaeta transversa Czerny, 1903; Fig. 53: left lateral; Fig. 54: posterior; Figs. 55–56: C. synneura (Hennig, 1938); Fig. 55: left lateral; Fig. 56: posterior; Figs. 57–58: C. pleuralis (Williston, 1896); Fig. 57: left lateral; Fig. 58: posterior.

FIGURES 61–66: Hypandrial complex, left lateral; Fig. 61: Craspedochaeta transversa Czerny, 1903; Fig. 62: C. chela spec. nov.; Fig. 63: C. pollostos spec. nov.; Fig. 64: C. pleuralis (Williston, 1896); Fig. 65: C. weemsi spec. nov.; Fig. 66: C. candida spec. nov.

FIGURES 71–77: Female internal genitalia (double line indicates the removal of an elongate median section to facilitate illustration); Fig. 71: Craspedochaeta feminea spec. nov.; Fig. 72: C. claripennis (Melander & Argo, 1924); Fig. 73: C. ornata Hennig, 1938; Fig. 74: C. transversa Czerny, 1903; Fig. 75: C. protomis spec. nov.; Fig. 76: C. nigrithorax (Hennig, 1938); Fig. 77: C. apsilutea spec. nov.