SCOLYTINI
publication ID |
https://doi.org/ 10.5281/zenodo.5178893 |
publication LSID |
lsid:zoobank.org:pub:0BB30FB2-E0A6-43AD-9476-E95DA74B605E |
persistent identifier |
https://treatment.plazi.org/id/039E87BC-B308-FFFA-FF01-F9A3287CF8C1 |
treatment provided by |
Felipe |
scientific name |
SCOLYTINI |
status |
|
TRIBE SCOLYTINI
Page 313
KEY: Wood 2007: 185 (to genera in South America).
(tx) Alonso-Zarazaga and Lyal 2009: 22. (ds, hb, tx) Wood 2007: 185.
Genus Camptocerus Dejean
Page 318
KEYS: Smith and Cognato 2010a: 16 (to all species); Wood 2007: 200 (to species in South America.
FIGURES: Smith and Cognato 2010a (various morphological features).
(an) López-Buenfil et al. 2001: 337; Smith and Cognato 2010a. (tx) Alonso-Zarazaga and Lyal 2009: 24. (ds, hb, tx) Smith and Cognato 2010a; Wood 2007: 199.
Page 318
aeneipennis (Fabricius) . Add:
FIGURES: Smith and Cognato 2010a: 62 (habitus, head), 85 (aedeagus); Wood 2007: PLATE 29.
DISTRIBUTION: SOUTH AMERICA: Bolivia.
(hb) Hulcr and Cognato 2010b. (ds, hb, tx) Smith and Cognato 2010a: 62; Wood 2007: 202.
Page 318
FIGURES: Smith and Cognato 2010a: 65 (habitus, head), 85 (aedeagus); Wood 2007: PLATE 30.
(ds, tx) Smith and Cognato 2010a: 64; Wood 2007: 204.
Page 318
FIGURES: Smith and Cognato 2010a: 66 (habitus, head), 85 (aedeagus); Wood 2007: PLATE 30.
DISTRIBUTION: SOUTH AMERICA: Ecuador, Peru.
(ds, tx) Smith and Cognato 2010a: 66; Wood 2007: 204.
Page 319
auricomus Blandford. Add C. striatulus Hagedorn as a synonym. Add:
FIGURES: López-Buenfil et al. 2001: 363 (proventriculus); Smith and Cognato 2010a: 32 (habitus, head), 85 (aedeagus); Wood 2007: PLATE 31.
DISTRIBUTION: NORTH AMERICA: Guatemala, Mexico (Veracruz). SOUTH AMERICA; Trinidad.
(ds, tx) Smith and Cognato 2010a: 32; Wood 2007: 207.
Page 319
costatus Chapuis. Add C. seriatus Eggers as a synonym. Add:
FIGURES: Smith and Cognato 2010a: 55,56 (habitus [see note below], head), 85 (aedeagus); Wood 2007: PLATE 31.
DISTRIBUTION: SOUTH AMERICA: Ecuador, French Guyana, Peru.
(hb) Smith and Cognato 2010a: 54. (ds, tx) Smith and Cognato 2010a: 54; Wood 2007: 205.
NOTE: Lateral and dorsal habitus figures of this species were omitted in Smith’s manuscript. Habitus figures listed above are actually of C. satyrus Smith and Cognato (Smith, personal communication)
Page 319
charpentierae Schedl. Remove from synonymy with C. cinctus Chapuis. Add :
FIGURES: Smith and Cognato 2010a: 42 (habitus, head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Brazil, Ecuador.
(ds, tx) Smith and Cognato 2010a: 42.
NOTE: Wood (2007) places this species in synonymy under C. cinctus Chapuis.
Page 320
cinctus Chapuis. Move View in CoL to C. suturalis (Fabricius) View in CoL as a synonym. Add: Synonymy by Smith and Cognato 2010a: 78.
(ds, tx) Wood 2007: 203.
Page 320
fasciatus (Fabricius) (= C. suturalis (Fabricius)) . Add:
Lectotype (male), Guyana, Essequibo River , designated by Smith and Cognato 2010a: 78. Page 318
gibbus (Fabricius) (= C. aeneipennis (Fabricus)) . Add:
Lectotype (female), Guyana, Essequibo River , designated by Smith and Cognato 2010a: 62.
Page 320
hirtipennis Schedl. Remove View in CoL from synonymy with C. suturalis (Fabricius) View in CoL . Add: FIGURES: Smith and Cognato 2010a: 70 (habitus, head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Brazil, Ecuador.
(ds, tx) Smith and Cognato 2010a: 69.
Page 319
infidelis Wood. Move View in CoL to C. opacicollis (Eggers) View in CoL as a synonym. Add: Synonymy by Smith and Cognato 2010a: 29.
(ds, tx) Wood 2007: 209.
Page 319
inoblitus (Schedl) . Add C. morio (Schedl) as a synonym. Add:
FIGURES: Smith and Cognato 2010a: 27 (habitus, head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Argentina.
HOST: Myrocarpus frondosus .
(ds) Bright 2004: 160. (ds, tx) Smith and Cognato 2010a: 26; Wood 2007: 208.
Page 319
FIGURES: Smith and Cognato 2010a: 49 (habitus, head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Ecuador.
(ds, tx) Smith and Cognato 2010a: 48; Wood 2007: 208.
Page 319
major (Eggers) . Add:
FIGURES: Smith and Cognato 2010a: 28 (habitus, head). (ds, tx) Smith and Cognato 2010a: 28; Wood 2007: 210.
Page 319
morio (Schedl) View in CoL . Move to C. inoblitus (Schedl) View in CoL as a synonym. Add: Synonymy by Smith and Cognato 2010a: 26.
(ds, tx) Wood 2007: 209.
Page 319
niger (Fabricius) . Add C. tectus Eggers as a synonym. Add:
FIGURES: Smith and Cognato 2010a: 35 (habitus, head), 85 (aedeagus); Wood 2007: PLATE 32.
DISTRIBUTION: SOUTH AMERICA: Bolivia, Ecuador, Peru, Venezuela.
(ds, tx) Smith and Cognato 2010a: 34; Wood 2007: 206.
Page 319
FIGURES: Smith and Cognato 2010a: 37 (habitus, head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Peru.
(ds, tx) Smith and Cognato 2010a: 37; Wood 2007: 208.
Page 319
opacicollis (Eggers) . Add C. infidelis Wood and C. uniseriatus Schedl as synonyms. Add:
FIGURES: Smith and Cognato 2010a: 29 (habitus, head), 85 (aedeagus); Wood 2007: PLATE 32.
DISTRIBUTION: NORTH AMERICA: Costa Rica. SOUTH AMERICA: Ecuador, Peru.
(ds, tx) Smith and Cognato 2010a: 29; Wood 2007: 209.
Page 320
FIGURES: Smith and Cognato 2010a: 39 (habitus, head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Bolivia.
(ds, tx) Smith and Cognato 2010a: 38; Wood 2007: 207.
Page 320
FIGURES: Smith and Cognato 2010a: 58, 59 (habitus, head), 85 (aedeagus); Wood 2007: PLATE 33.
(ds, tx) Smith and Cognato 2010a: 58; Wood 2007: 205.
Page 320
FIGURES: Smith and Cognato 2010a: 40 (habitus, head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Ecuador.
(ds, tx) Smith and Cognato 2010a: 40; Wood 2007: 207.
Page 320
seriatus Eggers. Move View in CoL to C. costatus Chapuis View in CoL as a synonym. Add: Synonymy by Wood 2007: 205.
Page 319
squammiger Chapuis (= C. niger (Fabricius)) Add :
Lectotype (male), French Guiana [Cayenne], designated by Smith and Cognato 2010a: 34. Page 319
striatulus Hagedorn. Remove from synonymy with C. niger (Fabricius) , move to C. auricomus Blandford as a synonym. Add:
Synonymy by Smith and Cognato 2010a: 32.
Page 320
suturalis (Fabricius) . Add C. cinctus Chapuis as a synonym. Add:
Lectotype (male), Guyana, Essequibo River , designated by Smith and Cognato 2010a: 78.
FIGURES: Hulcr and Cognato 2010b: 3209 (adults and gallery); Smith and Cognato 2010a: 79 (habitus, head), 85 (aedeagus); Wood 2007: PLATE 33 (specimen figured is C. hirtipennis Schedl (Smith, personal communication).
DISTRIBUTION: SOUTH AMERICA: French Guiana, Peru, Suriname, Venezuela.
HOST: Ocotea oblonga , Protium sp.
(hb) Hulcr and Cognato 2010b: 3209. (ds, tx) Wood 2007: 204. (ds, hb, tx) Smith and Cognato 2010a: 78.
Page 320
tectus Eggers. Remove from synonymy with C. orientalis , move to C. niger (Fabricius) as a synonym. Add:
Synonymy by Smith and Cognato 2010a: 34.
(ds, tx) Smith and Cognato 2010a: 34; Wood 2007: 206.
Page 319
uniseriatus Schedl. Remove from synonymy with C. major (Eggers) , move to C. opacicollis (Eggers) as a synonym. Add:
Synonymy by Smith and Cognato 2010a: 29.
ADD SPECIES:
annectans Wood 2007: 210. Holotype (female), BRAZIL: Serra de Baturite , Ceara; DEIM.
FIGURES: Smith and Cognato 2010a: 26 (habitus, head).
(tx) Smith and Cognato 2010a: 25.
coccoformus Smith and Cognato 2010a: 43 View in CoL . Holotype (male), BRAZIL: Amazonas, Hwy. ZF 2, km. 20.7, ca 60 km N of Manaus ; USNM.
FIGURES: Smith and Cognato 2010a: 44 (habitus, head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Brazil, Ecuador.
distinctus Smith and Cognato 2010a: 68 View in CoL . Holotype (male), ECUADOR: Napo, Reserva Ethnica Waorani GoogleMaps , 1 km S Onkone Gare Camp, 0.652778, 76.433333, 220 m; USNM.
FIGURES: Smith and Cognato 2010a: 68 (habitus, head), 85 (aedeagus).
doleae Smith and Cognato 2010a: 45 View in CoL . Holotype (male), ECUADOR: Napo, Tiputini Biodiversity Station GoogleMaps , 0.631944, 76.144167, 220-250 m; USNM.
FIGURES: Smith and Cognato 2010a: 46 (habitus, head), 85 (aedeagus). igniculus Smith and Cognato 2010a: 47 View in CoL . Holotype (male), BRAZIL: Amazonas, Hwy. ZF 2, km. 20.7, ca 60 km N of Manaus; USNM.
FIGURES: Smith and Cognato 2010a: 47 (habitus, head), 85 (aedeagus).
mallopterus Smith and Cognato 2010a: 50 View in CoL . Holotype (male), ECUADOR: Napo, Tiputini Biodiversity Station GoogleMaps , 0.631944, 76.144167, 220-250 m; USNM.
FIGURES: Smith and Cognato 2010a: 51 (habitus, head), 85 (aedeagus).
mandelshtami Petrov 2007: 101 View in CoL . Holotype (male), PERU: Loreto Province, 68 km SW in Iquitos to
Nauta , Itaya river, 4°11’S 73°26’W GoogleMaps ; ZMM.
FIGURES: Petrov 2007: 102 (habitus, antenna); Smith and Cognato 2010a: 72 (habitus,
head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Ecuador, French Guiana.
HOST: Protium sp.
(ds, tx) Smith and Cognato 2010a: 71.
noel Smith and Cognato 2010a: 73 View in CoL . Holotype (male), PERU: Loreto, 1.5 km N Teniente Lopez , 2°35.66’S
76°6.92’W, 210-240 m; SMUK.
FIGURES: Smith and Cognato 2010a: 74 (habitus, head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Bolivia, Brazil, Colombia, Ecuador, French Guiana,
Guyana, Peru, Suriname, Venezula.
HOSTS: Ocotea oblonga , Protium amazonicum .
petrovi Smith and Cognato 2010a: 52 View in CoL . Holotype (male), ECUADOR: Napo, Tiputini Biodiversity Station, 0.631944, 76.144167, 220-250 m; USNM. FIGURES: Smith and Cognato 2010a: 51 (habitus, head), 85 (aedeagus).
pilifrons Smith and Cognato 2010a: 56 View in CoL . Holotype (male), ECUADOR: Napo, Reserva Ethnica Waorani GoogleMaps , 1 km S Onkone Gare Camp, 0.652778, 76.433333, 220 m; USNM.
FIGURES: Smith and Cognato 2010a: 57 (habitus, head), 85 (aedeagus).
pseudoangustior Smith and Cognato 2010a: 76 View in CoL . Holotype (male), ECUADOR: Pastaza, Cononaco ; USNM.
FIGURES: Smith and Cognato 2010a: 76 (habitus, head), 85 (aedeagus).
DISTRIBUTION: SOUTH AMERICA: Brazil, Ecuador, Peru.
(hb) Smith and Cognato 2010a: 76.
satyrus Smith and Cognato 2010a: 53 View in CoL . Holotype (male), BRAZIL: Rondónia, 62 km SW Ariquemes , nr.
Fzda, Rancho Grande; FSCA.
FIGURES: Smith and Cognato 2010a: 55 (figures mislabeled as “ C. costatus ”), 85 (aedeagus). unicornis Smith and Cognato 2010a: 31 . Holotype (male), BRAZIL: Amazonas; USNM.
FIGURES: Smith and Cognato 2010a: 31 (habitus, head).
zucca Smith and Cognato 2010a: 60 View in CoL . Holotype (male), ECUADOR: Napo, Huahua Sumaco, km 44 on Hollin-Loreto Road ; CSCA.
FIGURES: Smith and Cognato 2010a: 60 (habitus, head).
Genus Cnemonyx Eichhoff
Page 313
KEY: Wood 2007: 186 (to all species in Central and South America).
(an) López-Buenfil et al. 2001: 336. (tx) Alonso-Zarazaga and Lyal 2009: 24; Rabaglia 2002: 801. (ds, tx) Wood 2007: 186.
Page 313
acuminatus Schedl. Add : (ds, tx) Wood 2007: 198.
Page 313
amazonicus (Eggers) View in CoL . Add: (ds, tx) Wood 2007: 192.
Page 314
barbatus Eggers. Move to C. galeritus Eichhoff View in CoL as a synonym. Add: Synonymy by Wood 2007: 199.
Page 314
boliviae (Blackman) . Add: ds, tx) Wood 2007: 194.
Page 314
Lectotype (female), Isla Martin Garcia , Buenos Aires, Argentina, designated by Wood 2007: 198.
(ds) Bright 2004: 160. (tx) Guadalupe del Rio, Lanteri and Suárez 2005: 4. (ds, tx) Wood 2007: 198.
Page 314
creber Schedl. Add View in CoL : (ds, tx) Wood 2007: 197.
Page 314
difformis (Schedl) View in CoL .Add: (ds, tx) Wood 2007: 196.
Page 314
(ds) Burgos-Solorio and Equihua 2007: 71.
Page 314
errans (Blandford) . Add:
(ds) Bright 2004: 160. (ds, tx) Wood 2007: 197.
Page 315
ficus (Schwarz) . Add:
(ds) Vázquez, Rodriguez and Zorrilla 2003: 18.
Page 315
flavicornis (Chapuis) . Add:
(ds) Bright 2004: 160. (ds, tx) Wood 2007: 190.
Page 315
furvescens Wood. Add View in CoL : (ds, tx) Wood 2007: 195.
Page 315
galeritus Eichhoff. Add View in CoL Minulus View in CoL barbatus Eggers as a synonym. Add: (ds, tx) Wood 2007: 199.
Page 316
glaber (Eggers) View in CoL . Add: (ds, tx) Wood 2007: 190.
Page 316
glabratus (Schedl) . Add:
FIGURE: López-Buenfil et al. 2001: 363 (proventriculus).
Page 316
hylurgoides (Schedl) View in CoL . Add: (ds, tx) Wood 2007: 192.
Page 316
insidiosus (Schedl) . Add:
(tx) Guadalupe del Rio, Lanteri and Suárez 2005: 6. (ds, tx) Wood 2007: 199.
Page 316
FIGURES: Wood 2007: PLATE 26. (ds, tx) Wood 2007: 194.
Page 316
jucundus Chapuis. Add : (ds, tx) Wood 2007: 193.
Page 316
longicollis (Blandford) View in CoL . Add: (ds, tx) Wood 2007: 198.
Page 316
maculicornis (Blandford) View in CoL . Add: FIGURES: Wood 2007: PLATE 27. (ds, tx) Wood 2007: 193.
Page 316
(ds, tx) Wood 2007: 196.
Page 317
niger (Eggers) View in CoL . Add: (ds, tx) Wood 2007: 194.
Page 317
panamensis (Blandford) . Add Loganius setulosus Eggers and L. similis Eggers as synonyms. Add:
FIGURES: Wood 2007: PLATE 27.
(ds, hb, tx) Wood 2007: 191.
Page 317
protivorus Wood. Add View in CoL : (ds, tx) Wood 2007: 192.
Page 317
rugulosus (Eggers) View in CoL . Add: FIGURES: Wood 2007: PLATE 28. (ds, tx) Wood 2007: 195.
Page 317
setulosus Eggers. Move View in CoL to C. panamensis (Blandford) View in CoL as a synonym. Add: Synonymy by Wood 2007: 191.
Page 317
similis Eggers. Move View in CoL to C. panamensis (Blandford) View in CoL as a synonym. Add: Synonymy by Wood 2007: 191.
Page 318
splendens (Wood) . Add:
(ds) Burgos-Solorio and Equihua 2007: 71.
Page 318
vagabundus (Wood) . Add:
(ds) Bright and Torres 2006: 394.
Page 318
vestitus (Eggers) View in CoL . Add: FIGURES: Wood 2007: PLATE 28. (ds, tx) Wood 2007: 190.
Page 315
vianai Schedl View in CoL (= C. flavicornis (Chapuis)) View in CoL . Add: (tx) Guadalupe del Rio, Lanteri and Suárez 2005: 8.
Page 318
vismiacolens Wood. Add View in CoL : FIGURES: Wood 2007: PLATE 29. (ds, tx) Wood 2007: 196.
Genus Scolytopsis Blandford View in CoL
Page 320
KEY: Wood 2007: 211 (to all species).
(an) López-Buenfil et al. 2001: 337. (tx) Alonso-Zarazaga and Lyal 2009: 24. (ds, hb, tx) Wood 2007: 210.
Page 321
argentinensis Eggers. Move View in CoL to S. toba Wichmann View in CoL as a synonym. Add: Synonymy by Wood 2007: 213.
(ds) Bright 2004: 160.
Page 320
brasiliensis Eggers. Add View in CoL : (ds, tx) Wood 2007: 212.
Page 321
bruchi Schedl. Move to S. toba Wichmann View in CoL as a synonym. Add: Synonymy by Wood 2007: 213.
(tx) Guadalupe del Rio, Lanteri and Suárez 2005: 5.
Page 321
cubensis Wood View in CoL (= S. puncticollis Blandford View in CoL ). Add: (ds) Vázquez, Rodriguez and Zorrilla 2003: 18.
Page 320
laticollis Wood. Add View in CoL : (ds, tx) Wood 2007: 212.
Page 320
orinocanus Wood. Add View in CoL : FIGURES: Wood 2007: PLATE 34. (ds, tx) Wood 2007: 213.
Page 320
peruanus Eggers. Add View in CoL : FIGURES: Wood 2007: PLATE 35. (ds, tx) Wood 2007: 211.
Page 321
FIGURE: López-Buenfil et al. 2001: 363 (proventriculus).
(ds) Burgos-Solorio and Equihua 2007: 71. (ds, tx) Wood 2007: 213.
Page 321
toba Wichmann. Remove from synonymy with S. puncticollis Blandford. Add S. argentinensis Eggers and S. bruchi Schedl as synonyms. Add:
(ds, tx) Wood 2007: 213.
Genus Scolytus Geoffroy
Page 321
KEYS: Furniss and Johnson 2002: 89 (to species in Idaho, USA); Mifsud and Knízek 2009: 35 (to species in Malta); LaBonte 2010: 209 (to some species in North America); de Laclos et al. 2004: 141-144 (to species in France); Mandelshtam and Petrov 2010a: 175 (to species in India); Petrov and Mandelshtam 2010: 99 (to species in Peru); Smith and Cognato 2010: 40 (to species in eastern North America); Wood 2007: 215 (to species in South America).
(an) López-Buenfil et al. 2001: 336. (tx) Alonso-Zarazaga and Lyal 2009: 24; Rabaglia 2002: 801. (ds, tx) Maiti and Saha 2009: 74. (hb) Faccoli 2000b. (tx, hb) Wood 2007: 214.
Page 322
DISTRIBUTION: ASIA: China (Hengduan Mtns.). (ds) Huang and Yin 1992: 853.
Page 355
abietis Blackman. Remove from synonymy with S. opacus . Add:
FIGURES: Equihua and Furniss 2009: 598 (pars stridens), 599 (abdominal sternites 2-5), 601 (gallery pattern).
(tx) Equihua-Martinez and Furniss 2009.
Page 322
FIGURES: Petrov and Mandelshtam 2010: 68 (male habitus), 69 (second sternite).
(tx) Petrov and Mandelshtam 2010: 67; Wood 2007: 226.
NOTE: In Wood 2007, descriptions of male and female are incorrect (Petrov and Mandelshtam 2010).
Page 322
FIGURES: Mifsud and Knízek 2009: 48 (habitus).
DISTRIBUTION: ASIA: Russia (Daghestan). EUROPE: Malta.
HOST: Prunus divaricata .
(ds) Bolu and Legalov 2008: 83; Colonnelli 2003: 97; Haack 2001: 274; Mifsud and Knízek 2009: 37; Petrov 2005: 219. (tx) Mandelshtam and Petrov 2010a: 175.
Page 324
angustatus Browne. Add S. facialis Schedl as a synonym. Add:
FIGURES: Petrov and Mandelshtam 2010: 70 (male frons), 71 (female frons). (tx) Petrov and Mandelshtam 2010: 69; Wood 2007: 227.
Page 324
FIGURES: Petrov and Mandelshtam 2010: 72 (habitus); Wood 2007: PLATES 35, 36.
DISTRIBUTION: SOUTH AMERICA: Peru.
(ds, tx) Petrov and Mandelshtam 2010: 71; Wood 2007: 224.
Page 324
atratus Chapuis. Move to S. carinatus Chapuis View in CoL as a synonym. Add: Synonymy by Wood 2007: 234.
Page 325
barbatus Schedl. Add : (tx) Wood 2007: 226.
Page 325
barinensis Wood. Add View in CoL : FIGURES: Wood 2007: PLATE 36. (hb, tx) Wood 2007: 233.
Page 325
belokanicus Stark. Move View in CoL to S. koenigi Schevyrew View in CoL as a synonym. Add: Synonymy by Petrov 2005: 220.
Page 325
FIGURE: Petrov and Mandelshtam 2010: 74 (habitus).
DISTRIBUTION: SOUTH AMERICA: Peru.
(ds, tx) Petrov and Mandelshtam 2010: 73: Wood 2007: 226.
Page 325
bicolor Eggers. Add View in CoL : (tx) Wood 2007: 223.
Page 325
binodus Wood. Add View in CoL : (tx) Wood 2007: 225.
Page 325
bolivianus Schedl. Add View in CoL : (tx) Wood 2007: 224.
Page 325
(ds) Mandelshtam and Petrov 2009a: 208.
Page 325
carinatus Chapuis. Add S. atratus Chapuis as a synonym. Add:
Lectotype (male), Cartagene, Bolivar, Colombia, designated by Wood 2007: 234. (tx) Wood 2007: 234.
Page 325 carpini (Ratzeburg) . Add: FIGURE: de Laclos et al. 2004: 154 (gallery pattern). DISTRIBUTION: ASIA: Russia (Daghestan). EUROPE: Denmark. (cn) Bocquillon 2002. (ds) Colonnelli 2003: 97; Mandelshtam, Nikitsky and Bibin 2005a: 23;
Noblecourt 2004: 33; Pedersen Runge and Jonsén 2008; Petrov 2005: 219; Voolma et al.
2004. (ds, hb) de Laclos et al. 2004: 154. Page 327
caudatus Eggers. Add S. conidens Browne as a synonym. Add: FIGURES: Wood 2007: PLATE 37.
(ds, tx) Wood 2007: 230.
Page 327
(tx) Mandelshtam and Petrov 2010a: 177. (ds, tx) Maiti and Saha 2009: 76.
Page 327
(ds) Mandelshtam and Petrov 2009a: 208.
Page 335
confusus (Eggers) . Add:
NOTE: Considered a valid species by Beaver and Liu 2010: 19, not synonym of S. japonicus Chapuis , original synonymy by Krivolutskaya, 1985 [see Wood and Bright Catalog, 1992].
Page 338
conidens Browne. Move View in CoL to S. caudatus Eggers View in CoL as a synonym. Add: Synonymy by Wood 2007: 230.
Page 328
convexus Schedl. Add View in CoL : (tx) Wood 2007: 224.
Page 328
costellatus Chapuis. Add S. pseudocostellatus Schedl and S. strigipennis Schedl as synonyms. Add:
FIGURES: Petrov and Mandelshtam 2010: 81 (male habitus); Wood 2007: PLATES 37, 38.
(tx) Petrov and Mandelshtam 2010: 78. (hb, tx) Wood 2007: 227.
Page 328
FIGURES: Wood 2007: PLATE 38.
(ds) Burgos-Solorio and Equihua 2007: 71. (tx) Petrov and Mandelshtam 2010: 80. (ds, hb, tx) Wood 2007: 230.
Page 328
FIGURES: Mandelshtam and Petrov 2010a: 173 (habitus), 174 (abdominal apex). (ds) Mandelshtam and Petrov 2009a: 208.
Page 329
FIGURES: Wood 2007: PLATE 39.
(ds) Vázquez, Rodriguez and Zorrilla 2003: 18. (ds, hb, tx) Wood 2007: 235.
Page 329
ecksteini Butovitsch. Add View in CoL : DISTRIBUTION: ASIA: Russia (Daghestan). (ds) Petrov 2005: 219.
Page 329
elongatus Schedl. Add View in CoL : (tx) Wood 2007: 223.
Page 329
ensifer Eichhoff. Add View in CoL : FIGURE: de Laclos et al. 2004: 146 (habitus). DISTRIBUTION: ASIA: Russia (Daghestan). EUROPE: Maltese Islands. (ds) Colonnelli 2003: 97; Mifsud and Colonnelli 2010; Noblecourt 2004: 33; Petrov 2005: 219;
Podlussány and György 2008: 199. (ds, hb) de Laclos et al. 2004: 146; Petrov and Nikitskii
2001: 171.
Page 331
facialis Schedl. Move to S. angustatus Browne as a synonym. Add: Synonymy by Petrov and Mandelshtam 2010: 69.
(tx) Wood 2007: 222.
Page 330
Neotype designated by Smith and Cognato 2010b: 37 “ Galesburg , Illinois ” (MCZ).
FIGURES: Smith and Cognato 2010b: 38 (male and female habitus, male and female frons, aedeagus); 39 (second sternite).
DISTRIBUTION: NORTH AMERICA: United States (Kansas, Ohio, Pennsylvania).
(ds, tx) Smith and Cognato 2010b: 35.
Page 331
frontalis Blandford. Add View in CoL : (tx) Beaver and Liu 2010: 18.
Page 331
golbachi Schedl. Add View in CoL : (ds, tx) Wood 2007: 229.
Page 331
intricatus (Ratzeburg) . Add:
DISTRIBUTION: ASIA: Russia (Daghestan, Leningrad, Yaroslavi). EUROPE: France ( Corsica).
(bv) Vrkocova et al. 2000. (ds) Colonnelli 2003: 97; de Laclos 2004: 166; Faccoli 2002: 100; Haack 2001: 274; Inglebert 2002: 124; Mandelshtam and Popovichev 2000: 891; Mandelshtam, Nikitsky and Bibin 2005a: 23; Petrov 2005: 219; Petrov and Nikitskii 2001: 171; Podlussány and György 2008: 199; Voolma et al. 2004. (hb) Petrov and Nikitskii 2001: 171. (tx) Vlasov 2005: 766. (ds, hb) de Laclos et al. 2004: 152.
Page 334
jacobsoni (Spessivtsev) . Add:
(ds) Mandelshtam and Petrov 2009a: 208.
Page 335
(ds) Mandelshtam and Petrov 2009a: 208. (tx) Beaver and Liu 2010: 19.
Page 335
jaroschevskyi Schevyrew. Add: DISTRIBUTION: ASIA: Russia (Daghestan). (ds) Petrov 2005: 220.
Page 336
(tx) Mandelshtam and Petrov 2010a: 177. (ds, tx) Maiti and Saha 2009: 77.
Page 336
FIGURES: Six et al. 2005: 230 (damage, galleries, habitus, diagnostic characters).
DISTRIBUTION: AFRICA: South Africa (introduced). ASIA: Russia (Daghestan). EUROPE: France ( Corsica), Switzerland. SOUTH AMERICA: Argentina, Brazil, introduced?.
HOSTS: Ulmus suberosa , U. laevis , U. foliacea .
(ds) Colonnelli 2003: 97; de Laclos 2004: 166; Germann 2010; Markovic 2000; Petrov 2005: 220. (ds, tx) Wood 2007: 221. (ds, ec, hb) Six et al. 2005.
NOTE: Treated as a subspecies in Colonnelli (2003) and Germann (2010).
Page 337
fasciatus Reitter (= S. kirschi Skalitzky ). Add:
(ds, tx) Colonnelli 2003: 97;
NOTE: Treated as a subspecies of S. kirschi Skalitzky in Colonnelli (2003);
Page 337
koenigi Schevyrew. Add S. belokanicus Stark as a synonym. Add: DISTRIBUTION: ASIA: Russia (Daghestan).
(ds) Colonnelli 2003: 97; Noblecourt 2004: 33. (ds, tx) Petrov 2005: 220.
Page 338
koltsei Reitter. Add:
(ds) Mandelshtam and Petrov 2009a: 208.
Page 339
laetus Wood. Add View in CoL : (tx) Wood 2007: 232.
Page 339 laevis Chapuis. Add View in CoL : FIGURE: de Laclos et al. 2004: 147 (ventral habitus). DISTRIBUTION: ASIA: Russia (Daghestan, Leningrad). HOSTS: Ulmus laevis View in CoL , U. suberosa View in CoL . (ds) Colonnelli 2003: 97; Mandelshtam and Popovichev 2000: 892; Mandelshtam, Nikitsky and Bibin 2005a: 24; Petrov 2005: 220; Petrov and Nikitskii 2001: 171; Voolma et al. 2004.
(hb) Petrov and Nikitskii 2001: 171. (ds, hb) de Laclos et al. 2004: 147. Page 340
(ds, hb, tx) Furniss and Johnson 2002: 90.
Page 340
(tx) Mandelshtam and Petrov 2010a: 175. (ds, tx) Maiti and Saha 2009: 78.
Page 341 mali (Bechstein) . Add: FIGURES: de Laclos et al. 2004: 151 (habitus); Mifsud and Knízek 2009: 48 (habitus). DISTRIBUTION: ASIA: Russia (Daghestan, Yaroslavi). NORTH AMERICA: United States
(Michigan). HOSTS: Cerasus sp. , Cedrus oblonga , Prunus armeniaca , P. divaricata , Mespilus germanica . (ds) Cognato et al. 2009: 110: Colonnelli 2003: 97; Haack 2001: 275; Mandelshtam, Nikitsky and Bibin 2005a: 24; Mifsud and Knízek 2009: 36; Petrov 2005: 220; Petrov and Nikitskii
2001: 171; Voolma et al. 2004. (hb) Petrov and Nikitskii 2001: 171. (tx) Vlasov 2005: 767. Page 344
marginatus Chapuis. Add : (tx) Wood 2007: 232.
Page 345
(ds, hb, tx) Furniss and Johnson 2002: 90.
Page 345
morawitzi Semenov. Add : DISTRIBUTION: ASIA: Russia (Yakutia).
(ds) Anonymous 2005a; Averensky and Yanovsky 2008: 26; CAB International 2007c; Faccoli 2002: 100: Vanhanen, Veteli and Niemelae 2008; Voolma et al. 2004.
Page 345
multistriatus (Marsham) . Add:
FIGURES: LaBonte 2010: 209 (various morphological features); de Laclos et al. 2004: 145 (gallery pattern); Lee et al. 2006 (habitus); López-Buenfil et al. 2001: 364 (proventriculus).
DISTRIBUTION: ASIA: Russia (Daghestan, Leningrad, Yaroslavi). NORTH AMERICA: Canada (Nova Scotia). SOUTH AMERICA: Argentina, Chile? (introduced, see note below).
(bv) McNee, Wood and Storer 2000; Seybold, Paine and Dreistadt 2008. (ds) Brockerhoff, Knízek and Bain 2003: 35; Colonnelli 2003: 97; Furniss and Johnson 2002: 91; Haack 2001: 275; Inglebert 2002: 124; Lee et al. 2009; Majka, Anderson and McCorquodale 2007: 418; Mandelshtam and Popovichev 2000: 892; Petrov 2005: 220; Petrov and Nikitskii 2001: 170; Voolma et al. 2004; Wood 2007: 222. (hb) Furniss and Johnson 2002: 91; Lee and Seybold 2010; Lee et al. 2010; Petrov and Nikitskii 2001: 170. (tx) Furniss and Johnson 2002: 91; Johnson et al. 2008; LaBonte 2010: 209; Vlasov 2005: 767; Wood 2007: 222. (bv, ec) Lee and Seybold 2010. (ds, hb) de Laclos et al. 2004: 145.
NOTE: Reports of introduction into South America are not confirmed. Species should not be included in South American fauna ( Wood 2007). Treated as a subspecies in Colonnelli (2003).
Page 354
FIGURE: López-Buenfil et al. 2001: 364 (proventriculus). Page 354
DISTRIBUTION: NORTH AMERICA: United States (Arkansas). (ds) Schiefer 2010: 111.
Page 354
(tx) Beaver and Liu 2010: 19.
Page 354
FIGURES: Petrov and Mandelshtam 2010: 89 (habitus); 90 (frons). (tx) Petrov and Mandelshtam 2010: 87; Wood 2007: 225.
NOTE: Holotype is a male.
Page 354
(tx) Mandelshtam and Petrov 2010a: 176. (ds, ec, hb) Buhroo and Ramamurthy 2008. (hb) Buhroo and Lakatos 2007; Buhroo, Chishti and Masoodi 2004.
Page 354
nodatus Wood. Add View in CoL : (hb, tx) Wood 2007: 228.
Page 355
nodulus (Wichmann) View in CoL . Add: (tx) Wood 2007: 235.
Page 355
novateutonicus Schedl. Add : FIGURES: Wood 2007: PLATE 40. (ds, tx) Wood 2007: 234.
Page 355
numidicus Brisout. Add : HOST: Cedrus libani atlantica . (ds) Benhalima et al. 2005.
Page 355
FIGURES: Equihua and Furniss 2009: 598 (pars stridens), 599 (abdominal sternites 2-5). (tx) Equihua-Martinez and Furniss 2009. (ds, hb, tx) Furniss and Johnson 2002: 92.
Page 353
orientalis Eggers (= S. multistriatus (Marsham) . Add: DISTRIBUTION: ASIA: Russia (Daghestan).
(ds) Petrov 2005: 220; Mandelshtam, Nikitsky and Bibin 2005a: 24. NOTE: Treated as a valid name by Knízek (2011).
Page 356
(tx) Petrov and Mandelshtam 2010: 88; Wood 2007: 233.
Page 356
piceae (Swaine) . Add:
FIGURES: LaBonte 2010: 210 (to various morphological features), 211 (to various features).
DISTRIBUTION: NORTH AMERICA: Canada (Nova Scotia)
(ds) Furniss, Holsten and Schultz 2002: 90; Majka, Anderson and McCorquodale 2007: 419. (tx) LaBonte 2010: 211. (ds, hb, tx) Furniss and Johnson 2002: 92.
Page 356
DISTRIBUTION: ASIA: China (Hengduan Mtns.). (ds) Huang and Yin 1992: 852.
Page 356
FIGURE: Petrov and Mandelshtam 2010: 91 (female habitus). (tx) Wood 2007: 228.
Page 357
DISTRIBUTION: ASIA: China (Hengduan Mtns.). (ds) Huang and Yin 1992: 853.
Page 357
DISTRIBUTION: NORTH AMERICA: United States (Wyoming [see Appendix]). (ds, hb, tx) Furniss and Johnson 2002: 93.
Page 357
(ds) Burgos-Solorio and Equihua 2007: 71. (ds, hb, tx) Wood 2007: 231.
Page 357
FIGURES: Petrov and Mandelshtam 2010: 92, 93 (male habitus); Wood 2007: PLATE 41.
DISTRIBUTION: SOUTH AMERICA: Peru.
(ds, tx) Petrov and Mandelshtam 2010: 90. (ds, hb, tx) Wood 2007: 232.
Page 357
pseudocostellatus Schedl. Move to S. costellatus Chapuis as a synonym. Add: Synonymy by Petrov and Mandelshtam 2010: 79.
(tx) Wood 2007: 227; Petrov and Mandelshtam 2010: 78.
Page 358 pygmaeus (Fabricius) . Add: FIGURE: de Laclos et al. 2004: 148 (habitus). DISTRIBUTION: ASIA: Russia (Daghestan, Yaroslavi). EUROPE: Great Britian. (ds) Colonnelli 2003: 97; Heal 2003a; Inglebert 2002: 124; Mandelshtam, Nikitsky and Bibin
2005a: 24; Markovic 2000; Petrov 2005: 220; Petrov and Nikitskii 2001: 170; Podlussány and György 2008: 199. (ec) Moser et al. 2005. (hb) Petrov and Nikitskii 2001: 170. (tx)
Vlasov 2005: 767. (ds, hb) de Laclos et al. 2004: 148. Page 361
DISTRIBUTION: ASIA: China (Hengduan Mtns.). (ds) Huang and Yin 1992: 853.
Page 361 ratzeburgi Janson. Add : DISTRIBUTION: ASIA: Russia (Altai, Daghestan, Yakutia, Yaroslavi). HOSTS: Betula pendula , B. litwinowi . (ds) Averensky and Yanovsky 2008: 26; Colonnelli 2003: 97; Demidko 2007: 34; Demidko 2007: 34; Haack 2001: 274; Mandelshtam and Petrov 2009a: 208; Mandelshtam and Popovichev 2000: 892; Mandelshtam, Nikitsky and Bibin 2005a: 24; Petrov 2005: 220; Petrov and Nikitskii 2001: 171; Voolma et al. 2004. (hb) Petrov and Nikitskii 2001: 171. (tx) Vlasov 2005: 767. (ds, hb) de Laclos et al. 2004: 149. Page 364
rugulosus (Müller) . Add:
FIGURES: de Laclos et al. 2004: 153 (habitus); Mifsud and Knízek 2009: 48 (habitus).
DISTRIBUTION: ASIA: Russia (Daghestan, Yaroslavi). EUROPE: Malta. NORTH AMERICA: Canada (Prince Edward Island).
HOSTS: Acer undulatum .
(ds) Ayberk and Cebeci 2010; Bolu and Legalov 2008: 83: Bright 2004: 160; Colonnelli 2003: 97; Foster 2011; Furniss and Johnson 2002: 93; Haack 2001: 275; Majka, Anderson and McCorquodale 2007: 419; Mandelshtam and Popovichev 2000: 892; Mandelshtam, Nikitsky and Bibin 2005a: 24; Mifsud and Knízek 2009: 37; Petrov 2005: 220; Petrov and Nikitskii 2001: 171; Podlussány and György 2008: 199; Voolma et al. 2004. (hb) Furniss and Johnson 2002: 93; Petrov and Nikitskii 2001: 171; Wood 2007: 221. (tx) Furniss and Johnson 2002: 93; Mandelshtam and Petrov 2010a: 175; Vlasov 2005: 767; Wood 2007: 221.
Page 369
mediterraneus Eggers View in CoL (= S. rugulosus (Müller) View in CoL . Add (ds) Al-Ahmadi and Salem 1999: 129.
Page 370
FIGURES: LaBonte 2010: 210, 212 (to various morphological features), 213 (lateral habitus); Lee et al. 2006 (habitus, gallery pattern, damage); Negrón et al. 2005: 84 (habitus), 88 (gallery pattern), 89 (damage).
DISTRIBUTION: ASIA: China (Hengduan Mtns.); Russia (Daghestan). NORTH AMERICA: Canada (Alberta, Manitoba, Ontario, Saskatchewan) ; United States (all western states), plus Connecticut, Minnesota, Missouri, Illinois, Indiana, Kansas, Maryland, Michigan, Nebraska, New Jersey, Ohio, Pennsylvania and Virginia [introduced] .
(bv) Lee and Seybold 2010; Seybold, Paine and Dreistadt 2008. (ds) CAB International 2009b; Cognato et al. 2009: 110; Haack 2006: 273; Lee et al. 2009; Mandelshtam and Petrov 2009a: 208; Petrov 2005: 220. (ec) Lee and Seybold 2010. (hb) Lee and Seybold 2010; Lee et al. 2010. (tx) Johnson et al. 2008; LaBonte 2010: 212. (ds, ec, hb) Lee et al. 2006, 2009, 2010; Negrón et al. 2005.
Page 371
scolytus (Fabricius) . Add:
FIGURE: de Laclos et al. 2004: 150 (epistomal margin).
DISTRIBUTION: ASIA: Russia (Daghestan).
(ds) Colonnelli 2003: 97; Faccoli 2004b; Haack 2001: 274; Inglebert 2002: 124; Mandelshtam, Nikitsky and Bibin 2005a: 24; Petrov 2005: 220; Podlussány and György 2008: 199; Voolma et al. 2004. (ec) Faccoli 2004b; Petrov and Nikitskii 2001: 170. (hb) Faccolim200b; Petrov and Nikitskii 2001: 170. (ds, hb) de Laclos et al. 2004: 150.
Page 376
semenovi (Spessivtsev) . Add:
(ds) Mandelshtam and Petrov 2009a: 209.
Page 371
seulensis Murayama View in CoL (= S. schevyrewi Semenov View in CoL ). Add: (ds) Huang and Yin 1992: 852.
Page 376
DISTRIBUTION: ASIA: China (Hengduan Mtns., Qinghai). (ds) Han et al. 2010; Huang and Yin 1992: 852.
Page 377
(tx) Wood 2007: 223.
Page 377
strigipennis Schedl. Move View in CoL to S. costellatus Schedl View in CoL as a synonym. Add: Synonymy by Petrov and Mandelshtam 2010: 79; Wood 2007: 227.
Page 377
submarginatus Schedl. Add : FIGURES: Wood 2007: PLATE 42. (ds, tx) Wood 2007: 234.
Page 377
(ds, hb, tx) Furniss and Johnson 2002: 93.
Page 377
FIGURES: Mifsud and Knízek 2009: 48.
DISTRIBUTION: ASIA: Russia (Daghestan). EUROPE: Malta.
(ds) Colonnelli 2003: 97; Mifsud and Knízek 2009: 35; Petrov 2005: 220. (ds, hb) Petrov and Nikitskii 2001: 170.
Page 378
Lectotype (male), Brazil, designated by Wood 2007: 236.
FIGURES: Wood 2007: PLATE 43.
(ds) Bright 2004: 161. (tx) Petrov and Mandelshtam 2010: 92; Wood 2007: 236. Page 378
torulus Wood. Add View in CoL : (tx) Wood 2007: 232.
Page 355
(ds) Colonnelli 2003: 97; Süda 2006;Voolma et al. 2004.
NOTE: Treated as a valid name in Colonnelli (2003) and Knízek (2011). Page 378
(ds) Mandelshtam and Petrov 2009a: 209.
Page 378
Page 379
tsugae (Swaine) . Add:
(ds, hb, tx) Furniss and Johnson 2002: 95.
FIGURES: LaBonte 2010: 210 (abdominal sternites). (ds, hb, tx) Furniss and Johnson 2002: 95.
Page 390
(bv) Seybold, Paine and Dreistadt 2008. (ec) Macias-Sámano and Borden 2000. (ds, hb, tx) Furniss and Johnson 2002: 95.
ADD SPECIES:
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
Kingdom |
|
Phylum |
|
Class |
|
Order |
|
Family |
SCOLYTINI
Bright, Donald E. 2014 |
cinctus
Smith, S. M. & A. I. Cognato 2010: 78 |
hirtipennis
Smith, S. M. & A. I. Cognato 2010: 70 |
infidelis
Smith, S. M. & A. I. Cognato 2010: 29 |
morio (Schedl)
Smith, S. M. & A. I. Cognato 2010: 26 |
coccoformus
Smith, S. M. & A. I. Cognato 2010: 43 |
distinctus
Smith, S. M. & A. I. Cognato 2010: 68 |
doleae
Smith, S. M. & A. I. Cognato 2010: 45 |
mallopterus
Smith, S. M. & A. I. Cognato 2010: 50 |
noel
Smith, S. M. & A. I. Cognato 2010: 73 |
petrovi
Smith, S. M. & A. I. Cognato 2010: 52 |
Smith, S. M. & A. I. Cognato 2010: 51 |
pilifrons
Smith, S. M. & A. I. Cognato 2010: 56 |
pseudoangustior
Smith, S. M. & A. I. Cognato 2010: 76 |
satyrus
Smith, S. M. & A. I. Cognato 2010: 53 |
zucca
Smith, S. M. & A. I. Cognato 2010: 60 |
frontalis
Beaver, R. A. & L. Y. Liu 2010: 18 |
seriatus
Wood, S. L. 2007: 205 |
acuminatus
Wood, S. L. 2007: 198 |
amazonicus (Eggers)
Wood, S. L. 2007: 192 |
barbatus
Wood, S. L. 2007: 199 |
boliviae (Blackman)
Wood, S. L. 2007: 194 |
creber
Wood, S. L. 2007: 197 |
difformis (Schedl)
Wood, S. L. 2007: 196 |
furvescens
Wood, S. L. 2007: 195 |
galeritus
Wood, S. L. 2007: 199 |
glaber (Eggers)
Wood, S. L. 2007: 190 |
hylurgoides (Schedl)
Wood, S. L. 2007: 192 |
jucundus
Wood, S. L. 2007: 193 |
longicollis (Blandford)
Wood, S. L. 2007: 198 |
maculicornis (Blandford)
Wood, S. L. 2007: 193 |
niger (Eggers)
Wood, S. L. 2007: 194 |
protivorus
Wood, S. L. 2007: 192 |
rugulosus (Eggers)
Wood, S. L. 2007: 195 |
setulosus
Wood, S. L. 2007: 191 |
similis
Wood, S. L. 2007: 191 |
vestitus (Eggers)
Wood, S. L. 2007: 190 |
vismiacolens
Wood, S. L. 2007: 196 |
argentinensis
Wood, S. L. 2007: 213 |
brasiliensis
Wood, S. L. 2007: 212 |
bruchi
Wood, S. L. 2007: 213 |
laticollis
Wood, S. L. 2007: 212 |
orinocanus
Wood, S. L. 2007: 213 |
peruanus
Wood, S. L. 2007: 211 |
atratus
Wood, S. L. 2007: 234 |
barbatus
Wood, S. L. 2007: 226 |
barinensis
Wood, S. L. 2007: 233 |
bicolor
Wood, S. L. 2007: 223 |
binodus
Wood, S. L. 2007: 225 |
bolivianus
Wood, S. L. 2007: 224 |
conidens
Wood, S. L. 2007: 230 |
convexus
Wood, S. L. 2007: 224 |
elongatus
Wood, S. L. 2007: 223 |
golbachi
Wood, S. L. 2007: 229 |
laetus
Wood, S. L. 2007: 232 |
marginatus
Wood, S. L. 2007: 232 |
nodatus
Wood, S. L. 2007: 228 |
nodulus (Wichmann)
Wood, S. L. 2007: 235 |
novateutonicus
Wood, S. L. 2007: 234 |
strigipennis
Wood, S. L. 2007: 227 |
submarginatus
Wood, S. L. 2007: 234 |
torulus
Wood, S. L. 2007: 232 |
vianai
Del Rio, M. G. & A. A. Lanteri & S. M. Suarez 2005: 8 |
belokanicus
Petrov, A. V. 2005: 220 |
ecksteini
Petrov, A. V. 2005: 219 |
ensifer
Petrov, A. V. 2005: 219 |
Laclos, E. & C. Mouy & L. Strenna & P. Agou 2004: 146 |
Colonnelli, E. 2003: 97 |
cubensis
Vazquez Moreno, L. L. & M. Rodriguez Perez & M. A. Zorrilla 2003: 18 |
mediterraneus
Al-Ahmadi, A. Z. & M Salem 1999: 129 |
seulensis
Huang, F. & H. Yin 1992: 852 |