Anisepyris columbianus ( Ashmead, 1893 )

Barbosa, Diego N. & Azevedo, Celso O., 2018, Revision of Anisepyris Kieffer (Hymenoptera, Bethylidae), with description of 135 new species, Zootaxa 4416 (1), pp. 1-258 : 95-96

Anisepyris columbianus ( Ashmead, 1893)

( Figs 13C ; 13D; 14J; 14N; 15A; 15D; 15K)

Epyris columbianus Ashmead, 1893 , 45: 60.

Rhabdepyris (Trichotepyris) columbianus ( Ashmead, 1893) . Combined by Kieffer, 1908, 76: 31. Rhabdepyris (Rhabdepyris) columbianus ( Ashmead, 1893) . Combined by Kieffer, 1914, 41: 357. Anisepyris pulchellus Fouts, 1928 , 30: 126. Synonym by Evans, 1959, 61: 104.

Rhabdepyris columbianus ( Ashmead, 1893) . Combined by Muesebeck & Walkley, 1951, 2: 729. Anisepyris columbianus ( Ashmead, 1893) . Combined by Evans, 1959, 61: 104.

Redescription, female. Body length 3.38 mm. Head and mesosoma black; wings clear hyaline. Head wide. Mandible with five distal teeth, wide, slightly angled, and with lower tooth longer than others. Median clypeal lobe rounded, short; median clypeal carina hardly distinct from clypeus. Frons coriaceous ; antennal scrobe carinate, but weak. Eye small. Frontal angle of ocellae forming right angle; ocelli very small. Vertex convex. Pronotal disc as long as wide, strongly coriaceous ; transverse pronotal carina complete; posterior margin with posterior pronotal sulcus. Notaulus narrow. Mesoscutellar disc with mesoscutellar sulcus narrow; posterior mesoscutellar sulcus present. Metapectal-propodeal disc long; metapostnotal median carina complete; longitudinal ridges present; metapostnotal-propodeal suture convergent to first pair of metapostnotal carina; paraspiracular carina present and conspicuous; metapleural carina present and conspicuous; metapectal-propodeal disc with short striae; propodeal declivity strigate. Mesopleuron with anterior fovea closed; mesopleural fovea closed; lower fovea divided; posterior fovea absent; mesopleural suture opened. Forewing with stigma developed; Rs&M vein not distinguished. Mesotibia not spinose; claws trifid. Genitalia. T9 with anterior arm long and overlapping 2vf.

Redescription, male. Body length 4.38 mm. Head and mesosoma black; wings clear hyaline; metasoma dark castaneous. Head wide. Flagellomeres short, with dense and short setae, with tooth projection at second and third flagellomere. Mandible narrow, curved, and with teeth equally wide. Median clypeal lobe quadrate, long, with apical median tooth. Frons coriaceous . Antennal scrobe carinate. Eye large. Frontal angle of ocellae forming right angle; ocelli small. Vertex convex. Pronotal disc as long as wide, strongly coriaceous ; transverse pronotal carina complete; with posterior pronotal sulcus. Notaulus narrow. Mesoscutellar sulcus narrow; posterior mesoscutellar sulcus present. Metapectal-propodeal disc short; metapostnotal median carina complete; longitudinal ridges present; metapostnotal-propodeal suture fused with first metapostnotal carina; paraspiracular carina present and conspicuous; metapleural carina present and conspicuous; metapectal-propodeal disc with short striae; propodeal declivity strigate. Mesopleuron with anterior fovea closed; mesopleural fovea closed; lower fovea divided; posterior fovea absent; mesopleural suture closed. Forewing with stigma developed; Rs&M vein not distinguished. Mesotibia not spinose. Claws trifid. Metasomal segments sparsely setose. Genitalia. Basiparamere shooter than paramere; paramere very wide, rounded, and with ventral and dorsal margin straight with short apical projection, sparsely setose; basivolsella with inner margin angled and with inner projection; cuspis with arms short, dorsal arm wide; aedeagus with basal portion angled and wide, and apical portion as long as basal and wide; apodeme slender.

Material examined. Lectotype ( E. columbianus Ashmead, 1893 ). Female. U.S.A., Washington D.C., 38°53'42"N 77°2'11"W, Type n° 14065 ( USNM). Paralectotype ( E. columbianus Ashmead, 1893 ). Male. [ U.S.A.], Fl[orid]a, Jacks[o]nville, Collection Ashmead ( USNM, #14065). U.S.A., Georgia, 1 ♂, Liberty Co[unty], S[ain]t Catherines Island, 19–23.vii.1990, Rozen, Quinter, Eickwort [collectors] ( AMNH); 2 ♂, 20–27.vi.1989, Rozen, Quinter, Foster &Krieger [collectors] ( AMNH); Arizona, 1 ♂, Cochise Co[unty], SWRS, 1.vii.1965, V. Roth [collector]( AMNH); Florida, 1 ♂, Alachua, Co[unty], Gainesville AEI, 29°35'53.6"N 82°21'54.8"W, May 2004, Malaise trap, D.B. Wahl ( AMNH); NICARAGUA, Granada, 1 ♂, Volcam Mombacho, San Joaquim 2, 16.iii.1998 —Mal[aise] Trap, J.M. Maes col[lector] ( SEAN); 2 ♂, El Progresso 1, 2.vi.1998; 2 ♂, El Progresso 2, 30.iv.1998; 2 ♂, San Joaquim 2, 30.iv.1998; 5 ♂, San Joaquim 3, 30.iv.1998; 2 ♂, 15.v.1998; 1 ♂, 21.vi.1998; 1 ♂, El Progresso 3, 15.v.1998; 4 ♂, Santa Ana 1, 15.v.1998; 8 ♂, Santa Ana 2, 15.v.1998.

Distribution. U.S. A: Washington, Florida, Georgia, Arizona, Florida; Mexico; Nicaragua; Costa Rica.

Figures

FIGURE 13. Anisepyris: columbianus species-group. Head in dorsal profile. A. A. arizonicus Evans, 1959; B. A. carinatus Santos & Azevedo, 2000; C. A. columbianus (Ashmead, 1893) (♀); D. A. columbianus (Ashmead, 1893) (♂); E. A. gibbosifrons Evans, 1959; F. A. grandis (Ashmead, 1887) (♀); G. A. grandis (Ashmead, 1887) (♂); H. A. insularis (Ashmead, 1894); I. A. interruptus Santos & Azevedo, 2000; J. A. laticeps Evans, 1959; K. A. nigripes Evans, 1966 (♀); L. A. nigripes Evans, 1966 (♂).