Incolacris C. Willemse, 1932

Storozhenko, Sergey Yu., 2021, On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae), Zootaxa 4970 (1), pp. 106-118 : 108

Genus Incolacris C. Willemse, 1932 , stat. resurr.

Incolacris C. Willemse, 1932: 150 (type species: Incolacris rubritarsi C. Willemse, 1932 , by original designation); C. Willemse, 1939: 89; Tinkham, 1940: 302; C. Willemse, 1956: 185.

Stolzia (part.): Hollis, 1975: 211; Otte, 1995: 338; Yin et al., 1996: 679; Li et al., 2006: 55.

Description. Head in frontal view elongated; frontal ridge broad, complete, almost reaching clypeus ( Fig. 5 ). Face in lateral view reclinated; frontal ridge almost straight ( Figs. 4, 8 ). Ventral genicular lobes of the hind femora triangular with distinct apical spine ( Fig. 12 ). Male 10th abdominal tergite without furculae ( Fig. 10 ). Male supra-anal plate triangular, with a median basal impression, lateral margins with two teeth in the basal half on each side or without it. Male cerci surpassing the supra-anal plate, compressed, conical, with pointed apex, without teeth on the inner side ( Fig. 11 ). Valves of ovipositor short, with apex rounded and strongly dentate ( Fig. 6 ). Male genitalia: epiphallus with broad bridge, ancorae present ( Fig. 15 ); the shape of other parts of genitalia unknown.

Species included. The genus consists of five species distributed in the Philippines, South China and Malaysia.

Remarks. Hollis (1975) synonymized Incolacris under Stolzia based the only on the asymmetrical shape of epiphallus but I disagree with such synonymy and consider it as a distinct genus differs from other genera of Incolacridinae in the specific shape of frontal ridge.

Figures

FIGURES 1–14. Subfamily Incolacridinae. 1. Bettotania cincifemur, female body, lateral view (schematic after Miller, 1935); 2. Stolzia rubromaculata, male head and pronotum, lateral view (schematic after C. Willemse, 1930); 3. Stolzia javana, female head, frontal view; 4–7. Incolacris rubritarsi, female: 4. Head and pronotum, lateral view; 5. Head, frontal view; 6. Apex of abdomen, lateral view (after Hollis, 1975); 7. The same, ventral view (after Hollis, 1975); 8–12. Incolacris jianfengensis comb. nov., male (after Zheng & Ma, 1989): 8. Head and pronotum, lateral view; 9. The same, dorsal view; 10. Apex of abdomen, dorsal view; 11. The same, lateral view; 12. Hind knee, lateral view; 13. Asymmetritania asymmetrica comb. nov., male apex of abdomen, dorsal view (after Ingrisch, 1989); 14. Asymmetritania vietnamensis comb. nov., male hind knee, lateral view.

FIGURES 15–24. Male genitalia of the subfamily Incolacridinae (15–18 after Hollis, 1975 and 19–24 after Ingrisch, 1989). 15. Incolacris trifasciata, epiphallus, dorsal view; 16. The same, Stolzia javana; 17. The same, Stolzia aberrans; 18. The same, Stolzia fasciata; 19–21. Bettotania maculata: 19. Epiphallus, dorsal view; 20. Phallic complex (epiphallus removed), dorsal view; 21. The same, lateral view; 22–24. Asymmetritania asymmetrica comb. nov.: 22. Epiphallus, dorsal view; 23. Phallic complex (epiphallus removed), lateral view; 24. The same, dorsal view. (Ae—ancorae; An—anterior projections of epiphallus; Ap—apical valves of penis; Apd—apodemes; Bp—basal valves of penis; Br—bridge; Cv—valves of cingulum; Fx—flexure; Lel—left lophi of epiphallus; Ler—right lophi of epiphallus; Os—oval sclerites; Rm—rami of cingulum; Sh—ectophallic sheath of penis; Zyg—zygoma).