Phthiracaridae Perty, 1841

Bolger, Thomas, 2017, Oribatid mites (Acari: Oribatida) recorded from Ireland: Catalogue, historical records, species habitats and geographical distribution, combinations, variations and synonyms, Zootaxa 4328 (1), pp. 1-174 : 28-37

Family Phthiracaridae Perty, 1841

Steganacarus (Atropacarus) Ewing, 1917

· Steganacarus (Atropacarus) striculus (C.L. Koch, 1835) *

· Original name: Hoplophora striculus C.L. Koch, 1835 .

· In: Deutschlands Crustaceen, Myriapoden und Arachniden, 2(10).

· Historical combinations/variations: Hoploderma striculum (C.L. Koch) Michael, 1898 ; Phthiracarus stricula (C.L. Koch) Oudemans, 1903 ; Hoplophora stricula (C.L. Koch) Ewing, 1917 ; Hoploderma stricula (C.L. Koch) Sellnick, 1928 ; Steganacarus striculus (C.L. Koch) Grandjean, 1933 ; Steganacarus striculum (C.L. Koch) Grandjean, 1933 .

· Synonyms: Steganacarus diaphanus Jacot, 1930 ; Steganacarus striculus diaphanus Jacot, 1938 ; Steganacarus senex Aoki, 1958 ; Steganacarus striculus insularis Weigmann , 1 976; Steganacarus trichosus Mahunka, 1982 .

Irish Reference/s and Habitat/s: Agricultural grassland (Curry 1969; Arroyo et al. 2015); cutaway raised bog ( Curry and Momen, 1988); ant nests ( Arroyo et al. 2015).

Counties: Kildare, Limerick and Offaly.

· Steganacarus (Atropacarus) wandae (Niedbała, 1981)

· Original name: Steganacarus wandae Niedbala, 1981 .

· In: Deux nouveaux Phthiracaridae ( Acari , Orbatidae) originaires de la Pologne. Bulletin de l'academie

polonaise des sciences serie des sciences biologiques, 28 (12), 699–766. · Historical combinations/variations: Atropacarus wandae (Niedbala) Arroyo et al. 2012 · Synonyms: Steganacarus navarrensis Moraza, 1985 .

Irish Reference/s and Habitat/s: Moss, bracken, soil on cliff and woodland (Bolger In press); moss on

soil surface in oak forest ( Arroyo et al. 2012, 2013b).

Counties: Laois and Mayo.

Hoplophthiracarus Jacot, 1933

· Hoplophthiracarus pavidus (Berlese, 1913)

· Original name: Hoploderma pavidus Berlese, 1913 .

· In: Acari nuovi. Manipoli VII-VIII. Redia , 9, 77–111.

· Historical com

· binations/variations: Calyptophthiracarus pavidus (Berlese) Balogh and Mahunka, 1983 ; Notophthiracarus (Calyptophthiracarus) pavidus (Berlese) Subías, 2004 .

· Synonyms: Hoplophthiracarus illinoisensis (Ewing, 1909) (?); Hoplophthiracarus cretensis Mahunka, 1979 ; Calyptophthiracarus cretensis inopinatus Mahunka, 1991 . Irish Reference/s and Habitat/s: Soil from Norway spruce forest ( Heneghan 1994). Counties: Kilkenny.

Phthiracarus (Phthiracarus) Perty, 1841

· Phthiracarus (Phthiracarus) affinis (Hull, 1914) *

· Original name: Hoploderma affinis Hull, 1914 .

· In: British Oribatidae . Notes on new and critical species. Naturalist, 1914, 215–288.

· Historical combinations/variations: Phthiracarus affinis (Hull) Turk, 1953 .

Irish Reference/s and Habitat/s: Forest soils ( Parry 1979; woodland, moss and ant nests (Bolger In press); pastures, broadleaved forestry (mixed stand of hawthorn and ash, mixed stand of hazel and beech), moss, soil and moss in canopy of oak forest, in Sitka spruce forest and blanket bogs ( Arroyo et al. 2009a); moss on soil in oak, ash, Scots pine and Sitka spruce forests, soil in Scots pine and Sitka spruce forests, canopy in Sitka spruce and moss in canopy in oak and Scots pine forests ( Arroyo et al. 2012, 2013b) moss in canopy and moss on soil in oak forest ( Arroyo et al. 2013c); riparian habitat (unpublished data); Atlantic blanket bogs ( Wisdom et al. 2011); agricultural grassland and ant nests ( Arroyo et al. 2015). Counties: Clare, Cork, Derry, Galway, Kerry, Kildare, Kilkenny, Laois, Limerick, Mayo, Offaly, Roscommon, Sligo, Tipperary, Waterford and Wicklow.

· Phthiracarus (Phthiracarus) dasypus (Dugès, 1834)

· Original name: Oribata dasypus Dugès, 1834 “sp. inq.”.

· In: Recherches sur l'ordre des Acariens. Troisième Mémoire. Annales des Sciences Naturelles (Zool), (2), vol. 2, 18–63.

· Historical combinations/variations: Ginglymacarus dasypus (Dugès) Ewing, 1917 , Hoploderma dasypus (Dugès) Halbert, 1907 .

· Synonyms: Hoplophora maxima Haupt, 1882 .

Irish Reference/s and Habitat/s: Moss ( Halbert 1907); Counties: Dublin.

· Phthiracarus (Phthiracarus) ferrugineus (C.L. Koch, 1841) *

· Original name: Hoplophora ferrugineus C.L. Koch, 1841 .

· In: Deutschlands Crustaceen, Myriapoden und Arachniden, Vol. 32 (10).

· Historical combinations/variations: Phthiracarus ferrugineus (C.L. Koch) Jacot, 1936 .

· Synonyms: Phthiracarus borealis Noordam and De Vlieger, 1943 ; Phthiracarus sellnicki Feider and Suciu, 1957 ; Phthiracarus ligneus Tarman, 1959 ; Phthiracarus juvenalis Parry, 1979 ; Phthiracarus membranifer Parry, 1979 ; Phthiracarus paraligneus Iturrondobeitia and Saloña, 1989 ; Archiphthiracarus ligneus Vasiliu, Ivan and Vasiliu, 1993 .

Irish Reference/s and Habitat/s: Soil from Norway spruce forest ( Heneghan 1994; Heneghan and Bolger, 1996; McCarthy 2004); fungal sporocarps ( O’Connell 1994; O’Connell and Bolger 1997; unpublished data).

Counties: Kilkenny and Wicklow.

· Phthiracarus (Phthiracarus) laevigatus (C.L. Koch, 1844)

· Original name: Hoplophora laevigatus C.L. Koch, 1844

· In: Deutschlands Crustaceen, Myriapoden und Arachniden, 38.

· Historical combinations/variations: Hoploderma laevigata (C.L. Koch) C.L. Koch, 1844 ; Hoploderma laevigatum (C.L. Koch) Tarman, 1955 ; Steganacarus laevigatus (C.L. Koch) Tarman, 1959 ; Pthiracarus nitens [sic] (C.L. Koch) Arroyo et al., 2013.

· Synonyms: Phthiracarus piger Scopoli, 1763 ; Phthiracarus contractilis Perty, 1841 ; Hoplophora nitens Nicolet, 1855 ; Phthiracarus roubali Berlese, 1923 ; Phthiracarus parabotrichus Feider and Sireticulacia, 1957 ; Archiphthiracarus parabotrichus (Feider and Sireticulacia) Vasiliu, Ivan and Vasiliu, 1993 ; Phthiracarus besuchetianus Mahunka and Mahunka-Papp, 2003 ; Phthiracarus nitens (Nicolet) Miko, 2016 . Irish Reference/s and Habitat/s: Fungal sporocarps ( O’Connell 1994); Sitka spruce forests and raised bogs ( Arroyo et al. 2009a); moss on soil in oak, ash, Scots pine and Sitka spruce forests ( Arroyo et al. 2012, 2013b); moss on soil in oak forest ( Arroyo et al. 2013c); riparian habitat (unpublished data); machair foredunes (Arroyo and Bolger 2011); agricultural grassland ( Arroyo et al. 2015); canopy in ash and Sitka spruce forests (unpublished data).

Counties: Clare, Kerry, Kildare, Limerick, Laois, Galway, Roscommon, Tipperary and Wicklow.

· Phthiracarus (Phthiracarus) longulus (C.L. Koch, 1841)

· Original name: Hoplophora longulus C.L. Koch, 1841 .

· In: Deutschlands Crustaceen, Myriapoden und Arachniden, Vol. 32 (17).

· Synonyms: Hoploderma borealis Trägårdh, 1910 ; Phthiracarus setosellus Jacot, 1929 ; Phthiracarus borealis (Trägårdh) Willmann 1931 [misidentification]; Phthiracarus prior Jacot, 1933 ; Phthiracarus montium Jacot, 1937 ; Phthiracarus apiculatus Jacot, 1939 ; Phthiracarus tardus Forsslund, 1943 ; Phthiracarus crenophilus Willmann, 1951 ; Phthiracarus pannonicus Willmann, 1954 ; Phthiracarus flexisetosus Parry, 1979 ; Phthiracarus hortobagyensis Mahunka, 1982 ; Phthiracarus mundus Niedbała, 1983 ; Phthiracarus malagensis Niedbała, 1986 .

Irish Reference/s and Habitat/s: Fungal sporocarps ( O’Connell 1994)

Counties: Kilkenny.

Phthiracarus (Archiphthiracarus) Balogh and Mahunka, 1979

· Phthiracarus (Archiphthiracarus) anonymus Grandjean, 1933 *

· Original name: Phthiracarus anonymus Grandjean, 1933 .

· In: Phthiracarus anonymum n. sp. Revue Française d’Entomologie, 1, 51–58.

· Historical combinations/variations: Paraphthiracarus anonymus (Grandjean) Aoki, 1980 ; Archiphthiracarus anonymus (Grandjean) Balogh and Mahunka, 1983 ; Pthiracarus anonymus [sic] (Grandjean) Arroyo and Bolger, 2011.

· Synonyms: Phthiracarus anonymus amicus Jacot, 1938 ; Archiphthiracarus pocsi Mahunka, 1983 ; Archiphthiracarus hauseri Mahunka, 1988 .

Irish Reference/s and Habitat/s: Forest soil fermentation layer ( Parry 1979), Salix ditch at seashore (Purvis 1982); fungal sporocarps ( O’Connell 1994; O’ Connell and Bolger 1997); soil from Norway spruce forest ( Heneghan 1994); mixed Norway spruce and Lodgepole pine forest ( Arroyo et al. 2009b); machair foredunes (Arroyo and Bolger 2011); moss in canopy of oak and Scots pine forests and soil and moss on soil in Sitka spruce forests ( Arroyo et al. 2012, 2013b); moss on trunk in yew forest and moss on soil in oak forest ( Arroyo et al. 2013c); canopy in ash forests (unpublished data).

Counties: Clare, Derry, Galway, Kerry, Kilkenny, Laois, Roscommon, Sligo, Wexford and Wicklow.

· Phthiracarus (Archipthiracarus) bryobius Jacot, 1930

· Original name: Phthiracarus setosellum bryobium Jacot, 1930 .

· In: Oribatid mites of the subfamily Phthiracaridae of the Northeastern United States. Proceedings of the Boston Society of Natural History, 39 (6), 209–261.

· Historical combinations/variations: Phthiracarus bryobius (Jacot) Jacot, 1938 ; Pthiracarus bryobius [sic] (Jacot) Arroyo and Bolger, 2011.

· Synonyms: Phthiracarus setosellum bryobium Jacot, 1930 ; Phthiracarus lanatus Tarman and Cervek 1976 ; Archiphthiracarus crinitosimilis Vasiliu, Ivan and Fabian 1994 ; Archiphthiracarus lanatus (Tarman and Cervek) sensu Vasiliu and Ivan 1995 .

Irish Reference/s and Habitat/s: Machair grassland (Arroyo and Bolger 2011); moss in canopy of Scots pine forest ( Arroyo et al. 2012, 2013b).

Counties: Mayo and Sligo.

· Phthiracarus (Archiphthiracarus) globosus (C.L. Koch, 1841)

· Original name: Hoplophora globosa C.L. Koch, 1841 .

· In: Deutschlands Crustaceen, Myriapoden und Arachniden, Vol. 32.

· Historical combinations/variations: Hoploderma globosum (C.L. Koch) Michael, 1898 ; Phthiracarus globosum (C.L. Koch) Willmann, 1931 ; Phthiracarus globosus (C.L. Koch) Willmann, 1931 .

· Synonyms: Phthiracarus rotundus Ewing, 1908 ; Phthiracarus subglobosus Berlese, 1923 ; Phthiracarus globus Parry, 1979 ; Phthiracarus mediocris Niedbała, 1984 .

Irish Reference/s and Habitat/s: Soil from Norway spruce forest ( Heneghan 1994). Counties: Kilkenny.

· Phthiracarus (Archiphthiracarus) montanus Pérez-Íñigo, 1969

· Original name: Phthiracarus montanus Pérez- Íñigo, 1969.

· In: Nuevos oribátidos de suelos españoles ( Acari, Oribatei ). EOS ,44, 377–403.

· Historical combinations/variations: Pthiracarus montanus [sic] (Pérez- Íñigo) Arroyo et al., 2013.

· Synonyms: Phthiracarus murphyi Harding, 1976 ; Phthiracarus rectisetosus Parry, 1979 ; Archiphthiracarus tzanoudakisi Mahunka, 1979 ; Phthiracarus assimilis Niedbała, 1983 ; Phthiracarus (Archiphthiracarus) tzanoudakisi Mahunka, 2008 .

Irish Reference/s and Habitat/s: Fungal sporocarps ( O’Connell 1994); soil from Norway spruce forest ( Heneghan 1994); machair grassland (Arroyo and Bolger 2011); moss on soil in ash, oak, Scots pine and Sitka spruce forests, moss in canopy in oak and Scots pine forests, soil in Sitka spruce forests and canopy in ash and Sitka spruce forests ( Arroyo et al. 2012, 2013b); moss in canopy in oak forest and moss on soil, moss on trunk, canopy and moss in canopy in yew forest ( Arroyo et al. 2013c); canopy in ash forests (unpublished data).

Counties: Clare, Donegal, Kerry, Kilkenny, Laois, Roscommon, Sligo and Wicklow.

· Phthiracarus (Archiphthiracarus) stramineus (C.L. Koch, 1841) “sp. inq.” *

· Original name: Hoplophora straminea Koch, 1841 .

· In: Deutschlands Crustaceen, Myriapoden und Arachniden, Vol. 32 (13).

· Historical combinations/variations: Phthiracarus stramineus (C.L. Koch) Jacot, 1936 ; Archiphthiracarus stramineus (C.L. Koch) Balogh and Mahunka, 1983 .

· Synonyms: Phthiracarus clavatus Parry, 1979 ; Phthiracarus opacus Niedbała, 1986 ; Phtiracarus clavatus [sic] (Parry) Arroyo et al., 2013.

Irish Reference/s and Habitat/s: Moss ( Halbert 1915); raised bogs ( Wisdom et al. 2011); moss on soil in oak forest and canopy of yew forest ( Arroyo et al. 2013c).

Counties: Dublin, Kerry, Mayo, Offaly and Roscommon.

Steganacarus (Steganacarus) Ewing, 1917

· Steganacarus (Steganacarus) magnus (Nicolet, 1855) * · Original name: Hoplophora magna Nicolet, 1855 .

· In: Histoire naturelle des Acariens qui se trouvent aux environs de Paris. Archives du Muséum d’Histoire Naturelle, Paris, 7, 381–482.

· Historical combinations/variations: Hoploderma magnum (Nicolet) Oudemans, 1900; Phthiracarus magna (Nicolet) Oudemans, 1915 ; Phthiracarus (Trachyhoplophora) magna (Nicolet) Berlese, 1923 ; Hoploderma magna (Nicolet) Sellnick, 1928 ; Steganacarus magnus (Nicolet) Willmann, 1941 .

· Synonyms: Phthiracarus magnus oblongus Oudemans, 1915 ; Phthiracarus magnus rotundus Oudemans, 1915 , nom. praeoc. by Ewing, 1908; Phthiracarus magnus transitans Oudemans, 1915 ; Steganacarus anomalus Sellnick, 1931 ; Steganacarus karamani Tarman, 1959 .

Irish Reference/s and Habitat/s: Moss, under bark and amongst fallen needles ( Halbert 1907, 1915); agricultural grassland ( Valarasan 1971), fungal sporocarps ( O’Connell 1994); fungal sporocarps ( O’Connell and Bolger 1997; unpublished data); bracken bed (Purvis 1982); grassland, heather and shelter belt on peat ( Longworth 1973); acid peat bog (Behan-Pelletier and Hill 1983); cutaway raised bog ( Curry and Momen 1988); bracken and woodland (Bolger In press); soil from Norway spruce forest, ( Heneghan 1994; Heneghan and Bolger 1996); raised bogs, mountain blanket bogs and Atlantic blanket bogs ( Wisdom et al. 2011); soil in oak, ash and Sitka spruce forests, moss on soil in ash, oak, Scots pine and Sitka spruce forest, moss in canopy in ash and Scots pine forests and canopy in oak and Scots pine forests ( Arroyo et al. 2012, 2013b); moss on soil and in trunks of oak and yew forest ( Arroyo et al. 2013c); ant nests ( Arroyo et al. 2015); canopy in ash forests (unpublished data).

Counties: Dublin, Clare, Cork, Kerry, Kilkenny, Laois, Limerick, Mayo, Offaly, Roscommon, Sligo, Tipperary, Wexford and Wicklow.

· Steganacarus (Steganacarus) michaeli Bernini and Avanzati, 1987

· Original name: Steganacarus michaeli Bernini and Avanzati, 1987 .

· In: Notulae Oribatologicae XLI. Some new Steganacarus species belonging to the magnus group (Acarida, Oribatida ). International Journal of Acarology, 13(3), 165–176.

Irish Reference/s and Habitat/s: Raised bogs and Atlantic blanket bos ( Wisdom et al. 2011); moss on soil in oak, ash, Scots pine and Sitka spruce forests, moss in canopy in oak, Scots pine and Sitka spruce forests, soil in oak and Scots pine forests and canopy in Scots pine forests ( Arroyo et al. 2012, 2013b); moss on soil and moss on trunks of oak and moss on soil in yew forest ( Arroyo et al. 2013c); riparian habitat (unpublished data); ant nests ( Arroyo et al. 2015); canopy in ash forests (unpublished data). Counties: Clare, Cork, Kerry, Kildare, Laois, Limerick, Mayo, Offaly, Roscommon, Sligo, Tipperary and Wicklow.