Pseudomanota perplexa, BLAGODEROV & GRIMALDI, 2004

BLAGODEROV, VLADIMIR & GRIMALDI, DAVID, 2004, Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae, American Museum Novitates 3433 (1), pp. 1-76 : 25-26

Pseudomanota perplexa , new species

Figures 28–30 , Plate 3D

DIAGNOSIS: As for the genus.

DESCRIPTION: Body length = 1.44 mm; wing length = 1.07 mm. Head: Eyes large, setulose, with large facets. Occiput and postgena setose, without erect bristles. Ocelli not seen. Antennae attached above middle of head. Frons, face and clypeus setose. Pedicel subconical. Flagellum 14­segmented, slightly thicker toward apex; flagellomeres cylindrical, width 1–1.5Χ length. Three palpomeres seen, basal one heart­shaped, penultimate one with long thin apical appendage about one­third its length, apical one long and slen­ der. Thorax: Antepronotum and proepisternum wide, completely divided, setose. Proepimeron narrow, rodlike. Scutum with lateral, acrostichal, and dorsocentral setae with bare strips in between. Anterior parapsidal suture distinct. Anepisternal suture declines posteriorly. Anepisternum with wide cleft. Metepisternum with long, narrow anterodorsal process touching laterotergite. Laterotergites and mediotergite bare. Wing membrane with macrotrichia and microtrichia. Humeral vein slightly distad of MA. Costa extends to R 5 apex at two­thirds distance between tips of R 5 and M 1, not reaching wing apex. Length of R 1 is 0.6Χ wing length. Small radial cell length 5Χ width. M 1 originates at apical one­third of wing, reaching wing margin before wing apex. M 2 originates at apical half of wing. M 3+4 free at the base. M 1, M 2, M 3+4, and CuA apices weak. Legs: Tibial setulae not arranged in rows except on apex of mid and hind tibiae. Tibial spurs ca. 3.5Χ tibia diameter. Hind tibia with a dorsal row of bristles. Abdomen setose, with 6 segments visible. Genital complex as long as 4th–6th segments combined, rotated 180°. Tergite 9 small, apex round, covering basal part of gonocoxites only. Aedeagus with two teeth apically. Gonocoxites separated, massive, setose. Gonostyli simple, sticklike, slightly curved, apically flat, shovel­like.

MATERIAL: Holotype AMNH Bu­599a, male. Myanmar: Katchin , from amber mines near Myitkyina.

ETYMOLOGY: The species epithet is from the Latin word perplexus, meaning ‘‘muddled, intricate’’, in reference to the intriguing combination of sciophiline and manotine characters.

TRIBE GNORISTINI EDWARDS, 1925

Figures

Figs. 28–30. Pseudomanota perplexa, n.sp. 28. Holotype AMNH Bu­599a. 29. Male genitalia dorsally. 30. Male genitalia ventrally.