Alavamanota Blagoderov and Arillo, 2002

BLAGODEROV, VLADIMIR & GRIMALDI, DAVID, 2004, Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae, American Museum Novitates 3433 (1), pp. 1-76 : 22-24

Alavamanota Blagoderov and Arillo, 2002

Alavamanota Blagoderov and Arillo, 2002: 34 .

DIAGNOSIS: Antepronotum and proepisternum completely divided; mediotergite and laterotergites bare; wing membrane with or without macrotrichia; R 1 relatively short, length of R 1 about the length of r­m; R 4 present; M 3+4 and CuA fork with stem.

TYPE SPECIES: Alavamanota hispanica Blagoderov and Arillo, 2002 .

COMMENTS: Closest to the recent genus Manota Williston, 1896 , distinguished by having the fourth palpomere attached preapically but very close to the apex of the third; two crossveins instead of one between R 1 and R 5 (RS1 and R 4); tibial trichia irregularly arranged; sternite 9 separate; sternite 8 without 4 strong protuberances bearing long setae; and the basal segment of each cercus small.

3+4

Alavamanota burmitina , new species

Figure 25 , Plate 3A

DIAGNOSIS: Flagellum compressed dorsoventrally; mesonotum with long lateral setae; wing membrane with macrotrichia; length of small radial cell 6Χ width; base of M 1 and M3 section weak; tibial setulae arranged in rows.

DESCRIPTION: Body length = 2.50 mm (holotype)/ 1.62–2.5 mm (paratypes); wing length = 1.69 mm (holotype)/ 1.08–1.85 mm (paratypes). Head: Postocciput with row of long protruding setae behind eye margin. Eyes slightly emarginate, densely setose, setae length 2Χ facet diameter. Facets round, densely set. Three ocelli in triangle, close to each other. Frons and face setulose. Antennae inserted above middle of head. Scape and pedicel subconical, with apical setae. Flagellum 14­segmented, flagellomeres compressed, widths 1.2–2Χ length, apical one conical. Texture of flagellomeres polygonlike. Only 3 segments of palpi visible, antepenultimate and penultimate with strong setae; penultimate 2.5Χ as long as preceding one, ovate; apical segment 1.7Χ length of penultimate one, narrow. Thorax: Scutum uniformly densely setose with short setae, bearing long lateral and posterior setae. Anterior parapsidal suture distinct. Suture between antepronotum and proepisternum complete, both segments uniform and covered with bristles. Scutellum with long setae. Anepisternum wider than higher, with wide dorsal cleft. Anepisternal suture declines posteriorly. Katepisternum setose ventrally. Mediotergite and laterotergites bare. Metepisternum with several light hairs posteroventrally. Wing membrane with macrotrichia; microtrichia not arranged in rows. Costa ends slightly beyond the tip of R 5, C and R 5 run very close to each other in apical part. R 1, R 5, r­m, M 1, and M 2 with setae ventrally and dorsally. Sc very short, ends free. Humeral vein oblique. R 1 about equal to length of r­m. Ratio of r­m and RS2 section is 1: 1.4. Base of M 1 and M 2 fork at the level of R 4. Small radial cell with length 6Χ width. Crossvein r­m horizontal, fused to tb, meets MA. M and CuA fork long, but not sessile. CuA curved caudally. Legs. Hind coxae bear apical only, not posterior setae. Tibial spur lengths ca. 4Χ tibial diameter. Tibial trichia in rows. Abdomen setose, with 6 visible segments. Gonocoxites fused, lighter then abdomen, setose, swollen. Gonostyli not seen.

MATERIAL: Holotype AMNH Bu­1271, male; paratypes AMNH Bu­428a, Bu 279a, males. Myanmar: Katchin, from amber mines near Myitkyina.

SUBFAMILY SCIOPHILINAE WINNERTZ, 1863 TRIBE SCIOPHILINI WINNERTZ, 1863 Neuratelia Rondani, 1856

Neuratelia Rondani, 1856: 195 .

Anaclinia Winnertz, 1863:770 .

Proanaclinia Meunier, 1904:145 .

Odontopoda Aldrich, 1897: 187 .

DIAGNOSIS: As given by Vockeroth (1972): Tibiae with distinct bristles, length of tibial spurs twice the tibia diameter; anepisternum, mesepimeron, and metepisternum bare; wing membrane with macrotrichia; sternite 8 of male large; gonocoxites partly or completely fused ventrally, gonostyli complex, subdivid­ ed or with elaborate processes.

TYPE SPECIES: Mycetophila nemoralis Meigen, 1818: 256 (orig. des.).

COMMENTS: Two species of the genus were described from Baltic amber ( Meunier, 1904)

Neuratelia maimecha , new species

Figure 26 , Plate 3B

DIAGNOSIS: Sc short, ends before RS base; length of RS1 section 2.5Χ that of crossvein r­m; base of M 3+4 and CuA fork at the middle of r­m; mediotergite with sparse, fine trichia.

DESCRIPTION: Body length = 2.11 mm (rest); wing length = 2.05 mm. Head: Flagellum 14­segmented, flagellomeres cylindrical, with length about equal to width. Scape and pedicel rounded. Mouthparts form proboscis slightly shorter than head height. Palpi 4­segmented, basal and antepenultimate segments oval, penultimate and apical segment bacilliform, apical slightly longer and narrower than penultimate, length ratio 1:3:4: 5.5. Thorax: Scutum with numerous, short, scattered setae and long lateral ones. Antepronotum with three setae, proepisternum with five. Anepisternum bare, with deep cleft in posterior part. Anepisternal suture declines posteriorly. Metepisternum quadrate, touching katepisternum. Laterotergites with several setae, mediotergite with sparse, fine trichia caudally. Wing membrane with microtrichia and few macrotrichia. Costa ends beyond tip of R 5. Sc meets C just before RS base. Sc 2 absent. Longitudinal veins with setae. M 3+4 and CuA base slightly distad of M3 base. Legs: Tibiae with distinct bristles. Fore tibia longer than first tarsomere.

MATERIAL: Holotype PIN 3311 /661, end of abdomen not preserved, sex unknown. Russia: Taimyr Peninsula , Yantardakh, coll. 1971.

ETYMOLOGY: Species epithet is derived from the Maimecha river on the Taimyr Pen­ insula, Siberia, where the amber deposit is located.

Figures

Figs. 25, 26. Alavamanota and Neuratelia. 25. A. burmitina, n.sp., holotype AMNH Bu­1271. 26. N. maimecha, n.sp., holotype PIN 3311/661.