Xylographellina Kawanabe & Miyatake, 1996

Lopes-Andrade, Cristiano, 2008, An essay on the tribe Xylographellini (Coleoptera: Tenebrionoidea: Ciidae), Zootaxa 1832 (1), pp. 1-110 : 10

publication ID	https://doi.org/10.11646/zootaxa.1832.1.1
publication LSID	lsid:zoobank.org:pub:45A82AD6-F369-4A74-A280-048BAE6E2DA5
persistent identifier	https://treatment.plazi.org/id/038E87B5-FFBA-CE79-FF6A-4AF8FC57FCF5
treatment provided by	Felipe (2021-07-23 11:52:45, last updated by Plazi 2023-11-03 12:38:44)
scientific name	Xylographellina Kawanabe & Miyatake, 1996
status

Treatment

Subtribe Xylographellina Kawanabe & Miyatake, 1996 ( Figs 1–6 View FIGURES 1–4 View FIGURES 5–8 , 9–10 View FIGURES 9–12 , 13–135 View FIGURES 13–15 View FIGURES 16–18 View FIGURES 19–20 View FIGURES 21–22 View FIGURES 23–25 View FIGURE 26 View FIGURES 27–29 View FIGURES 30–32 View FIGURES 33–35 View FIGURES 36–38 View FIGURES 39–41 View FIGURES 42–44 View FIGURES 45–47 View FIGURES 48–50 View FIGURES 51–53 View FIGURES 54–56 View FIGURES 57–59 View FIGURES 60–62 View FIGURES 63–65 View FIGURES 66–68 View FIGURES 69–71 View FIGURES 72–73 View FIGURES 74–76 View FIGURES 77–79 View FIGURE 80 View FIGURE 81 View FIGURES 82–84 View FIGURES 85–87 View FIGURES 88–90 View FIGURES 91–93 View FIGURES 94–95 View FIGURES 96–97 View FIGURES 98–100 View FIGURE 101 View FIGURES 102–104 View FIGURES 105–107 View FIGURES 108–110 View FIGURES 111–113 View FIGURES 114–116 View FIGURES 117–119 View FIGURES 120–122 View FIGURES 123–125 View FIGURES 126–127 View FIGURES 128–129 View FIGURES 130–133 View FIGURE 134 View FIGURE 135 , 159 View FIGURE 159 )

Included genera

Xylographella Miyatake, 1985 (type genus)

Scolytocis Blair, 1928

Diagnosis

This subtribe is characterized by the combination of the following characters: (i) scutellum conspicuous, its basal width more than 0.10X the greatest elytral width; (ii) prosternal process laminate; (iii) procoxae subconical, extending below the plane of the prosternal process; (iv) apex of prementum acute; (v) female terminalia devoid of gonostyli.

Distribution

Xylographellina is the most widespread subtribe of Xylographellini ( Figs 26 View FIGURE 26 , 80–81 View FIGURE 80 View FIGURE 81 , 101 View FIGURE 101 , 134 View FIGURE 134 , 159 View FIGURE 159 ), occurring throughout the Neotropical Region and in parts of the Palearctic (restricted to Japan), Oceanic, Australasian (restricted to New Zealand) and Indo-Malayan Regions.

References

Blair, K. G. (1928) Heteromera, Bostrychoidea, Malacodermata and Buprestidae. In: Insects of Samoa and Other Samoan Terrestrial Arthropoda, Part 45. Coleoptera. Fasc. 2. British Museum (Natural History), London, 67 - 109.

Kawanabe, M. & Miyatake, M. (1996) A redescription of Xylographella punctata (Coleoptera, Ciidae), with description of a new tribe. Elytra, Tokyo, 24 (1), 125 - 130.

Miyatake, M. (1985) Ciidae, pp. 278 - 285. IN: Y. Kurasawa, S. Hisamatsu and H. Sasaji (eds.), The Coleoptera of Japan in Color, 3. Hoikusha, Osaka (in Japanese).

Figures

FIGURE 26. Distribution map of Xylographella Miyatake, showing the known distribution of Xyl. punctata Miyatake (triangles) in Japan and Xyl. speciosa sp. nov. (stars) in the Philippines.

FIGURE 80. Distribution map of the continental Northern Neotropical species of Scolytocis Blair, showing the known distributions of Scol. bouchardi sp. nov. (stars), Scol. danielssoni sp. nov. (double circles), Scol. difficillimus sp. nov. (fulfilled square), Scol. howdeni sp. nov. (open squares), Scol. indecisus sp. nov. (plus symbols), Scol. lawrencei sp. nov. (fulfilled triangles), Scol. panamensis sp. nov. (open circle).

FIGURE 81. Distribution map of the insular Northern Neotropical species of Scolytocis Blair, showing the known distributions of Scol. cariborum sp. nov. (plus symbol) and Scolytocis kiskeyensis sp. nov. (stars).

FIGURE 101. Map showing the distribution of the Southern Neotropical species of Scolytocis. Scol. fritzplaumanni sp. nov. (circles), Scol. furieriae sp. nov. (triangles), Scol. paschoali sp. nov. (squares).

FIGURE 134. Distributional map of the Indo-Malayan species of Scolytocis Blair, showing the distribution of Scol. malayanus sp. nov. (triangles), Scol. philippinensis sp. nov. (double circle), Scol. thayerae sp. nov. (letters “X”) and Scol. werneri sp. nov. (plus symbol).

FIGURE 159. Distribution map of Xylographellini, showing the known distributions of the subtribes Xylographellina Kawanabe & Miyatake (stars) and Syncosmetina subtrib. nov. (triangles).

FIGURES 1–4. Scolytocis paschoali sp. nov. 1. Part of the head, frontal view showing the eye. 2. Antennal club, showing the sensillifers (arrows). Note that the club is compact, with antennomeres not freely articulated. 3. Head, ventral view showing the mouthparts. Note the longitudinal sulcus of prementum (arrow). 4. Ventral view showing the metaventrite.

FIGURES 5–8. Prementum (pm) with labial palpi (l.p). Note that the apical labial palpomeres are enlarged (small arrows), and that the apex of prementum is acute in Xylographella and Scolytocis, and rounded in Syncosmetus and Tropicis (see big arrows). 5. Xyl. punctata Miyatake, lateral view. 6. Scol. paschoali sp. nov. 7. Sync. reticulatus Miyatake. 8. Trop. sexcarinatus (Waterhouse).

FIGURES 9–12. Female terminalia, showing the gonostyli (gs), gonocoxites (gc), baculi of basal gonocoxites (small arrows), paraprocts (pp), baculi of paraprocts (b.pp), proctiger (pt), and spiculum ventrale (sv). The baculi of proctiger are barely sclerotized, so they are not indicated. 9. Xylographella punctata Miyatake. 10. Scolytocis furieriae sp. nov. 11. Syncosmetus japonicus Sharp. 12. Tropicis sexcarinatus (Waterhouse).

FIGURES 13–15. Xylographella punctata Miyatake. 13. Dorsal view. 14. Lateral view. 15. Ventralview.

FIGURES 16–18. Xylographella speciosa sp. nov. 16. Dorsal view. 17. Lateral view. 18. Ventral view.

FIGURES 19–20. Hindwings of the two species of Xylographella Miyatake. 19. Xyl. punctata Miyatake. 20. Xyl. speciosa sp. nov.

FIGURES 21–22. Xylographella Miyatake. 21. Antennae of Xyl. punctata Miyatake (A) and Xyl. speciosa sp. nov. (B). Note that the first antennomere is not shown in the latter (asterisk). 22. Metatibiae of Xyl. punctata Miyatake (A) and Xyl. speciosa sp. nov. (B).

FIGURES 23–25. Xylographella Miyatake. 23. Tegmen (teg) and penis (pen) of Xyl. punctata. 24. Y-Shaped ninth segment (broken, see asterisks) of Xyl. punctata. 25. Male genitalia of Xyl. speciosa sp. nov. showing the Y-shaped ninthsegment (ix), tegmen (teg) and penis (pen).

FIGURE 26. Distribution map of Xylographella Miyatake, showing the known distribution of Xyl. punctata Miyatake (triangles) in Japan and Xyl. speciosa sp. nov. (stars) in the Philippines.

FIGURES 27–29. Scolytocis bouchardi sp. nov. 27. Dorsal view. 28. Lateral view. 29. Ventral view.

FIGURES 30–32. Scolytocis cariborum sp. nov. 30. Dorsal view. 31. Lateral view. 32. Ventral view.

FIGURES 33–35. Scolytocis danielssoni sp. nov. 33. Dorsal view. 34. Lateral view. 35. Ventral view.

FIGURES 36–38. Scolytocis difficillimus sp. nov, holotype. 36. Dorsal view. 37. Lateral view. 38. Ventral view.

FIGURES 39–41. Scolytocis howdeni sp. nov. 39. Dorsal view. 40. Lateral view. 41. Ventral view.

FIGURES 42–44. Scolytocis indecisus sp. nov. 42. Dorsal view. 43. Lateral view. 44. Ventral view.

FIGURES 45–47. Scolytocis kiskeyensis sp. nov, holotype. 45. Dorsal view. 46. Lateral view. 47. Ventral view.

FIGURES 48–50. Scolytocis lawrencei sp. nov, holotype. 48. Dorsal view. 49. Lateral view. 50. Ventral view.

FIGURES 51–53. Scolytocis panamensis sp. nov. 51. Dorsal view. 52. Lateral view. 53. Ventral view.

FIGURES 54–56. Hindwings of Northern Neotropical Scolytocis Blair. 54. Scol. bouchardi sp. nov. 55. Scol. cariborum sp. nov. 56. Scol. danielssoni sp. nov.

FIGURES 57–59. Hindwings of Northern Neotropical Scolytocis Blair. 57. Scol. difficillimus sp. nov. 58. Scol. howdeni sp. nov. 59. Scol. indecisus sp. nov.

FIGURES 60–62. Hindwings of Northern Neotropical Scolytocis Blair. 60. Scol. kiskeyensis sp. nov. 61. Scol. lawrencei sp. nov. 62. Scol. panamensis sp. nov.

FIGURES 63–65. Antennae of the Northern Neotropical Scolytocis Blair. 63. Scol. bouchardi sp. nov. (A), Scol. cariborum sp. nov. (B), Scol. danielssoni sp. nov. (C). 64. Scol. difficillimus sp. nov. (A), Scol. howdeni sp. nov. (B), Scol. indecisus sp. nov. (C). 65. Scol. kiskeyensis sp. nov. (A), Scol. lawrencei sp. nov. (B), Scol. panamensis sp. nov. (C). First antennomere not shown in Scol. danielssoni sp. nov. (63C, asterisk).

FIGURES 66–68. Metatibiae of the Northern Neotropical Scolytocis Blair. 66. Scol. bouchardi sp. nov. (A), Scol. cariborum sp. nov. (B), Scol. danielssoni sp. nov. (C). 67. Scol. difficillimus sp. nov. (A), Scol. howdeni sp. nov. (B), Scol. indecisus sp. nov. (C). 68. Scol. kiskeyensis sp. nov. (A), Scol. lawrencei sp. nov. (B), Scol. panamensis sp. nov. (C).

FIGURES 69–71. Male genitalia of Scolytocis Blair showing the Y-shaped ninth segment (ix), basal piece (b.p), tegmen (teg) and penis (pen). 69. Scol. bouchardi sp. nov. 70. Scol. cariborum sp. nov. (ninth segment broken; see asterisks). 71. Scol. difficillimus sp. nov.

FIGURES 72–73. Male genitalia of Scolytocis Blair showing the Y-shaped ninth segment (ix), tegmen (teg) and penis (pen). 72. Scol. howdeni sp. nov. 73. Scol. indecisus sp. nov. The dashed line indicates the membranous basal portion of tegmen. Note that the apical portion of tegmen is extremely sclerotized.

FIGURES 74–76. Male genitalia of Scolytocis kiskeyensis sp. nov. showing the Y-shaped ninth segment (ix), tegmen (teg) and penis (pen). 74. Tegmen and penis of specimen from the Dominican Republic (type locality). 75. Tegmen and penis of specimen from Puerto Rico. 76. Ninth segment (pregenital segment) of specimen from Puerto Rico.

FIGURES 77–79. Male genitalia of Scolytocis Blair showing the Y-shaped ninth segment (ix), basal piece (b.p), tegmen (teg) and penis (pen). 77. Scol. lawrencei sp. nov., specimen from Costa Rica. The dashed line indicates the membranous basal portion of tegmen. 78. Scol. lawrencei sp. nov., specimen from Panama. 79. Scol. panamensis sp. nov.

FIGURES 82–84. Scolytocis fritzplaumanni sp. nov, holotype. Note the presence of small cuticular globules (protuberances) on the head, pronotum and elytra (small arrows). 82. Dorsal view. 83. Lateral view. 84. Ventral view.

FIGURES 85–87. Scolytocis furieriae sp. nov, holotype. 85. Dorsal view. 86. Lateral view. 87. Ventral view.

FIGURES 88–90. Scolytocis paschoali sp. nov, holotype. 88. Dorsal view. 89. Lateral view. 90. Ventralview.

FIGURES 91–93. Hindwings of the Southern Neotropical Scolytocis Blair. 91. Scol. fritzplaumanni sp. nov. 92. Scol. furieriae sp. nov. 93. Scol. paschoali sp. nov.

FIGURES 94–95. Southern Neotropical Scolytocis Blair. 94. Antennae of Scol. fritzplaumanni sp. nov. (A), Scol. furieriae sp. nov. (B) and Scol. paschoali sp. nov. (C). 95. Metatibiae of Scol. fritzplaumanni sp. nov. (A), Scol. furieriae sp. nov. (B) and Scol. paschoali sp. nov. (C). First antennomere not shown in Scol. furieriae sp. nov. (94B, asterisk).

FIGURES 96–97. Male genitalia of Scolytocis fritzplaumanni sp. nov. showing the Y-shaped ninth segment (ix), basal piece (b.p), tegmen (teg) and penis (pen). 96. Specimen from Nova Teutônia, Santa Catarina, Brazil (type locality). 97. Specimen from Guaratuba, Rio Grande do Sul, Brazil.

FIGURES 98–100. Male genitalia of Scolytocis Blair showing the Y-shaped ninth segment (ix), basal piece (b.p), tegmen (teg) and penis (pen). 98. Scol. furieriae sp. nov. from Jussari, Bahia, Brazil (type locality). 99–100. Scol. paschoali sp. nov., specimens from Macaé de Cima, Rio de Janeiro, Brazil (type locality). 99. Dorsal-ventral view. 100. Lateral view.

FIGURES 102–104. Scolytocis malayanus sp. nov., holotype. 102. Dorsal view. 103. Lateral view. 104. Ventral view.

FIGURES 105–107. Scolytocis novaezelandiae sp. nov., holotype. 105. Dorsal view. 106. Lateral view. 107. Ventral view.

FIGURES 108–110. Scolytocis philippinensis sp. nov., holotype. 108. Dorsal view. 109. Lateral view. 110. Ventral view.

FIGURES 111–113. Scolytocis thayerae sp. nov., holotype. 111. Dorsal view. 112. Lateral view. 113. Ventralview.

FIGURES 114–116. Scolytocis werneri sp. nov., holotype. 114. Dorsal view. 115. Lateral view. 116. Ventral view.

FIGURES 117–119. Scolytocis zimmermani sp. nov., holotype. 117. Dorsal view. 118. Lateral view. 119. Ventral view.

FIGURES 120–122. Hindwings of the new Indo-Malayan (in part) and Australasian Scolytocis Blair. 120. Scol. malayanus sp. nov. 121. Scol. novaezelandiae sp. nov. 122. Scol. philippinensis sp. nov.

FIGURES 123–125. Hindwings of the new Indo-Malayan (in part) and Oceanic Scolytocis Blair. 123. Scol. thayerae sp. nov. 124. Scol. werneri sp. nov. 125. Scol. zimmermani sp. nov.

FIGURES 126–127. Antennae of the Indo-Malayan, Australasian and Oceanic species of Scolytocis Blair. 126. Scol. malayanus sp. nov. (A), Scol. novaezelandiae sp. nov. (B) and Scol. philippinensis sp. nov. (C). 127. Scol. thayerae sp. nov. (A), Scol. werneri sp. nov. (B) and Scol. zimmermani sp. nov. (C).

FIGURES 128–129. Metatibiae of the Indo-Malayan, Australasian and Oceanic species of Scolytocis Blair. 128. Scol. malayanus sp. nov. (A), Scol. novaezelandiae sp. nov. (B) and Scol. philippinensis sp. nov. (C). 129. Scol. thayerae sp. nov. (A), Scol. werneri sp. nov. (B) and Scol. zimmermani sp. nov. (C).

FIGURES 130–133. Male genitalia of the Indo-Malayan species of Scolytocis Blair showing the Y-shaped ninth segment (ix), tegmen (teg) and penis (pen). 130. Scol. malayanus sp. nov. 131. Scol. philippinensis sp. nov. 132. Scol. thayerae sp. nov. 133. Scol. werneri sp. nov.

FIGURE 135. Distributional map of the Australasian and Oceanic species of Scolytocis Blair, showing the distribution of Scol. novaezelandiae sp. nov. (triangles), Scol. samoensis Blair (plus symbol) and Scol. zimmermani sp. nov. (square).