Pugnatrypaea, Poore & Dworschak & Robles & Mantelatto & Felder, 2019

Poore, Gary C. B., Dworschak, Peter C., Robles, Rafael, Mantelatto, Fernando L. & Felder, Darryl L., 2019, A new classification of Callianassidae and related families (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support, Memoirs of Museum Victoria (Mem. Mus. Vic.) 78, pp. 73-146 : 97-98

Pugnatrypaea gen. nov.

http://zoobank.org/urn:lsid:zoobank.org:act:0F41845B-F526-

4AB8-80B7-A60B6538919E

Type species. Callianassa pugnatrix De Man, 1905 , by present designation.

Diagnosis. Rostrum acute, anteriorly directed, as long as eyestalks. Pleomere 1 tergite fused, divided into 2 sections by transverse step. Cornea with scattered reduced pigmentation. Antennular peduncle from about as long exceeded by all of antennal peduncular article 5; articles 2 and 3 with single lateral row of 6–10 well-spaced long setae along lower margin.

Maxilliped 3 merus distally oblique with obtuse angle between distal and lower margins, without distal spine on mesial margin. Male major cheliped merus with simple proximal hook on lower margin; propodus distal margin with deep notch at base of fixed finger. Pereopod 3 propodus oval, lower margin slightly convex, leading to narrow sharply rounded proximal lobe. Male pleopod 2 present or absent. Uropodal endopod ovoid, usually longer than wide, anterior margin straight or slightly convex, posterodistal margin evenly convex, with spiniform setae near anterior and distal margins, or with facial spiniform setae on rib. Uropodal exopod about 1.5–1.8 times as long as wide, distal margin clearly differentiated from anterior margin, anterodistal corner right-angled, posterodistal margin with row of 6–8 long blade-like setae proximal to long setae on distal margin. Telson anterolateral lobe obsolete, undefined; tapering over distal third to pair of posterior lobes separated by deep notch, with medial spine.

Etymology. An alliteration of the name of the type species and Trypaea , a genus of the family.

Remarks. The telson tapers over its distal third to a pair of lobes separated by a deep notch with a medial spine. The two pairs of spiniform setae typically found on the posterolateral angles of the telson of callianassids are prominently enlarged and displaced anteriorly. Pugnatrypaea pugnatrix and an unidentified species Pugnatrypaea GMX lie on the same molecular clade (Robles et al. 2019). Both have a simple curved spine on the lower margin of the merus of the major cheliped. The antennular peduncle is shorter than the antennal peduncle in Pugnatrypaea GMX (fig. 6j) while they appear to be of similar lengths in Pugnatrypaea pugnatrix (De Man, 1928: fig. 23). Species with similar maxillipeds 3, telsons and short antennular peduncles are P. intermedia and P. lobetobensis but these differ in having a truncate, instead of rounded, uropodal exopod and the merus of the major cheliped dentate along the lower margin and with a spine on the upper margin.