Laccophilus tonkinensis Brancucci, 1983

Hájek, Jiří & Brancucci, Michel, 2015, A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae), Raffles Bulletin of Zoology 63, pp. 309-326 : 324-325

Laccophilus tonkinensis Brancucci, 1983

( Figs. 16 , 34 )

Laccophilus tonkinensis Brancucci, 1983: 258 (original description); Hájek, 2003: 120 (new records).

Type locality. Tonkin, Hoa Binh [= Vietnam, Hoa Binh Province] .

Material examined. 17 specimens. CHINA: Hainan: 4 ex., Limushan Mts. , exposed pools along road, partly connected with stream, 19°9.1–2′N, 109°45–46′E, 550–750 m, coll. M. Fikáček & S. Zhao ( NMPC, ZSMG) ; 1 male, Jianfengling Mts., Tianchi Lake env., rd from Tianchi vill. To “sector 5”, pools in shaded drying-up stream in primary forest, 18°43.6–44.1′N, 108°52.1–5′E, 820–950 m, 10 May 2011, coll. M. Fikáček & S. Zhao ( NMPC) . LAOS: Attapeu Province: 3 males, road No. 18, km 80 E Attapeu, stream near the road, 17°48.6′N, 107°23.4′E, ca. 665 m, 29 April 2010, coll. J. Hájek ( NMPC) . Khammouan Province: 1 male 4 females, Nakai-Nam Theun NPA, Ban Navang env., 17°57–59′N, 105°13–16′E, 600–750 m, 18–21 May 2012, NHMB Basel Expedition Laos 2012, M. Brancucci, M. Geiser, K. Phanthavong, S. Xayalath ( NHMB) . V IETNAM: 4 ex., Tonkin , coll. de Cooman ( IRSNB) .

Diagnosis. Medium sized, elongate oval species (body length 3.8–4.1 mm). Elytra with subbasal testaceous band broad, more or less connected with elytral base ( Fig. 16 ). Apical part of median lobe in lateral view with distinct bulge on dorsal side; apex elongate with pointed tip ( Fig. 34 ).

Habitat. In Hainan, Laccophilus tonkinensis was collected in pools in shaded drying up stream in primary mountane forest. In southern Laos, it was collected in a pool in a small forest stream with a clay bottom and a layer of decaying leaves ( Fig. 39 ).

Distribution. A species described from northern Vietnam and subsequently recorded from Hainan Island, China ( Brancucci, 1983; Hájek, 2003). Here we present the first records from central and southern Laos ( Fig. 40 ).

Figures

Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm.

Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm.

Figs. 36–39. Habitats: 36, E Kalimantan, Muara Ritan (type locality of Laccophilus kalimantanensis, new species); 37, China, Guangxi, Shiwandashan (locality of L. komareki, new species); 38, E Kalimantan, Sungai Wain (locality of L. mahakamensis); 39, Laos, Attapeu (locality of L. tonkinensis).

Figs. 40–41. Distribution of the species of the Laccophilus javanicus group