Luthela Xu & Li, 2022

Xu, Xin, Yu, Li, Liu, Fengxiang & Li, Daiqin, 2022, Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae), Zootaxa 5091 (1), pp. 131-154 : 134-135

publication ID	https://doi.org/10.11646/zootaxa.5091.1.5
publication LSID	lsid:zoobank.org:pub:7E744DAE-E9E6-4FB5-9897-1CCF03D373BC
DOI	https://doi.org/10.5281/zenodo.5843597
persistent identifier	https://treatment.plazi.org/id/C6E2A81A-368B-4799-AC41-782E4C33BE16
taxon LSID	lsid:zoobank.org:act:C6E2A81A-368B-4799-AC41-782E4C33BE16
treatment provided by	Plazi (2022-01-12 11:31:14, last updated 2024-11-29 13:16:44)
scientific name	Luthela Xu & Li
status	gen. nov.

Treatment

Genus Luthela Xu & Li , gen. nov.

Type species: Luthela yiyuan sp. nov.

Diagnosis. Males of Luthela can be distinguished from those of Songthela Ono, 2000 by the contrategular margin having regular larger teeth ( Figs 3D View FIGURE 3 , 5D View FIGURE 5 , 6D View FIGURE 6 , 7D View FIGURE 7 , 10H View FIGURE 10 , 12D View FIGURE 12 , 14D View FIGURE 14 ); from those of the other genera of Heptathelinae by the conductor with a smooth surface, and with two long spines, upper and lower ones ( Figs 3D View FIGURE 3 , 5D View FIGURE 5 , 6D View FIGURE 6 , 7D View FIGURE 7 , 10H View FIGURE 10 , 12D View FIGURE 12 , 14D View FIGURE 14 ). Luthela females differ from those of Songthela by the middle receptacular clusters having shorter stalks ( Figs 4 View FIGURE 4 , 5H View FIGURE 5 , 6H–J View FIGURE 6 , 8 View FIGURE 8 , 9 View FIGURE 9 , 11 View FIGURE 11 , 13 View FIGURE 13 , 14H–M View FIGURE 14 ); from those of the other genera of Heptathelinae by the positions of two pairs of the receptacular clusters: middle ones situated at anterior margin of bursa copulatrix and lateral ones situated at dorsolateral position of bursa copulatrix ( Figs 4 View FIGURE 4 , 5H View FIGURE 5 , 6H–J View FIGURE 6 , 8 View FIGURE 8 , 9 View FIGURE 9 , 11 View FIGURE 11 , 13 View FIGURE 13 , 14H–M View FIGURE 14 ).

Description. Male body length, 12.5–18.6 (N = 18), female, 12.5–27.0 (N = 85); male palpal conductor with a smooth surface, and with two long spines, upper and lower ones ( Figs 3D View FIGURE 3 , 5D View FIGURE 5 , 6D View FIGURE 6 , 7D View FIGURE 7 , 10H View FIGURE 10 , 12D View FIGURE 12 , 14D View FIGURE 14 ); contrategular margin with regular teeth ( Figs 3D View FIGURE 3 , 5D View FIGURE 5 , 6D View FIGURE 6 , 7D View FIGURE 7 , 10H View FIGURE 10 , 12D View FIGURE 12 , 14D View FIGURE 14 ); tegulum with three distinct apophyses ( Figs 3C View FIGURE 3 , 5F View FIGURE 5 , 6E View FIGURE 6 , 7F View FIGURE 7 , 10F View FIGURE 10 , 12F View FIGURE 12 , 14F View FIGURE 14 ); female genitalia with two paired receptacular clusters, middle ones situated at anterior margin of bursa copulatrix with tubular stalks, and lateral ones situated more dorsolaterally ( Figs 4 View FIGURE 4 , 5H View FIGURE 5 , 6H–J View FIGURE 6 , 8 View FIGURE 8 , 9 View FIGURE 9 , 11 View FIGURE 11 , 13 View FIGURE 13 , 14H–M View FIGURE 14 ).

Etymology. The genus name is created by a combination of ‘Lu-’, referring to Shandong Province, China, and ‘thela’, referring to spinnerets as the Greek word thele means nipple-like protuberance ( Ono 2000). The gender is feminine.

Composition. L. yiyuan sp. nov. (type species), L. luotianensis ( Yin et al., 2002) , comb. nov., L. schensiensis ( Schenkel, 1953) , comb. nov., L. badong sp. nov., L. dengfeng sp. nov., L. handan sp. nov., L. taian sp. nov., L. yuncheng sp. nov.

Distribution. China (Hebei, Henan, Hubei, Shandong, Shaanxi, and Shanxi Provinces).

References

Ono, H. (2000) Zoogeographic and taxonomic notes on spiders of the subfamily Heptathelinae (Araneae, Mesothelae, Liphistiidae). Memoirs of the National Science Museum, 33, 145 - 151.

Schenkel, E. (1953) Chinesische Arachnoidea aus dem Museum Hoangho-Peiho in Tientsin. Boletim do Museu Nacional do Rio de Janeiro, 119, 1 - 108.

Yin, C. M., Tang, G., Zhao, J. Z. & Chen, J. (2002) Two new species of the genus Heptathela from China (Araneae: Liphistiidae). Acta Arachnologica Sinica, 11, 18 - 21.

Figures

FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm.

FIGURE 4. Female genital anatomy of Luthela yiyuan sp. nov. A–E, K–O, vulva dorsal view. F–J, P–T, vulva ventral view. A, F, XUX-2012-034. B, G, XUX-2012-035. C, H, XUX-2012-049. D, I, XUX-2016-101. E, J, XUX-2016-107. K, P, XUX-2012- 042. L, Q, XUX-2012-040. M, R, XUX-2012-046. N, S, XUX-2016-103. O, T, XUX-2016-108. Scale bars: 0.5 mm.

FIGURE 5. Male and female genital anatomy of Luthela badong sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. (H) vulva dorsal view. I, vulva ventral view. A–G, XUX-2012-140 (holotype). H, I, XUX-2012-141. Scale bars: 0.5 mm.

FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. H–J, vulva dorsal view. K–M, vulva ventral view. A–G, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm.

FIGURE 7. Male genital anatomy of Luthela handan sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2011-214 (holotype). D–G, XUX-2011-209. Scale bars: 0.5 mm.

FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm.

FIGURE 9. Female genital anatomy of Luthela luotianensis (Yin et al., 2002) comb. nov. A, B, vulva dorsal view. C, D, vulva ventral view. A, C, XUX-2012-080. B, D, XUX-2012-082. Scale bars: 0.5 mm.

FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm.

FIGURE 11. Female genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm.

FIGURE 12. Male genital anatomy of Luthela taian sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2014-143A (holotype). D–G, XUX-2014-139. Scale bars: 0.5 mm.

FIGURE 13. Female genital anatomy of Luthela taian sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2014-133. B, E, XUX-2014-135. C, F, XUX-2014-143. G, J, XUX-2014-144. H, K, XUX-2014-151. I, L, XUX-2014- 154. Scale bars: 0.5 mm.

FIGURE 14. Male and female genital anatomy of Luthela yuncheng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. H–J, vulva dorsal view. K–M, vulva ventral view. A–G, XUX-2011-235 (holotype). H, K, XUX-2011-230. I, L, XUX-2011-234. J, M, XUX-2011-229. Scale bars: 0.5 mm.