Heterhelus, Jacquelin du Val, 1858

Hisamatsu, Sadatomo, 2011, A review of the Japanese Kateretidae fauna (Coleoptera: Cucujoidea), Acta Entomologica Musei Nationalis Pragae 51 (2), pp. 551-585 : 563

publication ID	https://doi.org/10.5281/zenodo.5329313
DOI	https://doi.org/10.5281/zenodo.5397257
persistent identifier	https://treatment.plazi.org/id/0388D148-EA06-8730-FEC5-FE0AFBD76719
treatment provided by	Felipe (2021-08-28 15:03:19, last updated by Plazi 2023-11-05 07:37:42)
scientific name	Heterhelus
status

Treatment

Key to subgenera and species of Japanese Heterhelus View in CoL View at ENA

1 Labrum shallowly notched. Antenna with distinct 3-segmented club ( Fig. 8E View Fig ). Pronotum strongly convex. Parameres of male genitalia abruptly bent inward at mid-length ( Fig. 8A View Fig ). Head dark brown, always darker than elytra. Subgenus Boreades Parsons, 1943 View in CoL . .. ............................................................................................... H. (B.) solani ( Heer, 1841)

– Labrum deeply notched. Antenna with indistinct 3-segmented club ( Figs. 6O View Fig , 7D View Fig ). Pronotum feebly convex. Parameres of male genitalia regularly arcuate inwards ( Figs. 6A, 6D View Fig , 7A, 7E View Fig ). Head always the same color as elytra. Subgenus Heterhelus Jacquelin du Val, 1858 View in CoL ......................................................................................................................... 2

2 Body uniformly black; sides of pronotum regularly arcuate, never angulate; parameres of male genitalia short and wide (LP/WP = 2.1 (n = 1)) ( Figs. 7A, 7E View Fig ); apex of median lobe bluntly rounded in lateral aspect ( Figs. 7C, 7F View Fig ); base of ovipositor pigmented ( Fig. 7G View Fig ). ........................................................................................... H. (H.) morio ( Reitter, 1878)

– Body reddish-yellow to blackish; sides of pronotum regularly arcuate or strongly angulate; parameres of male genitalia long and slender (LP/WP = 3.1 (n = 1)) ( Figs. 6A, 6D View Fig ); apex of median lobe sharply acuminate in lateral aspect ( Figs. 6C, 6E View Fig ); base of ovipositor not pigmented ( Fig. 6F View Fig ). ...................................................... H. (H.) scutellaris ( Heer, 1841)

References

HEER O. 1841: Fauna Coleopterorum Helvetica. Pars 1., fasciculus 3. Impensis Orelii, Fuesslini et Sociorum, Turici, 361 - 652 pp.

PARSONS C. T. 1943: A revision of Nearctic Nitidulidae (Coleoptera). Bulletin of the Museum of Comparative Zoology 92: 121 - 278.

REITTER E. 1878: Coleopterorum species novae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch Botanischen Gesellschaft in Wien 27: 165 - 194.

Figures

Fig. 8. Heterhelus (Boreades) solani (Heer, 1841). A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – ovipositor; E – right antenna, male; F – pronotum; G – left maxillary palpus.A–G = Hokkaido, Kawayu.

Fig. 6. Heterhelus (Heterhelus) scutellaris (Heer, 1841).A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – parameres of the lectotype of H. japonicus; E – median lobe of the lectotype of H. japonicus (lateral view); F – ovipositor; G – male abdominal sternite VII; H–N – variation of pronotum (H – Hokkaido; I – Aomori; J – Niigata; K – Tokyo; L – Yamaguchi; M – Ehime; N – Fukuoka); O – male right antenna. A, B, C, G, O = Hokkaido, Hitsujigaoka; F = Ehime Pref., Omogokei.

Fig. 7. Heterhelus (Heterhelus) morio (Reitter, 1878). A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – male right antenna; E – parameres of the lectotype of H. morio; F – median lobe of the lectotype of H. morio (lateral view); G – ovipositor; H – male abdominal sternite VII; I – pronotum. A, B, C, D, G, H, I = Ehime Pref., Omogokei.