Heptapactes, Schmidt, 2007

Schmidt, Christian, 2007, Revision of the Neotropical Scleropactidae (Crustacea: Oniscidea), Zoological Journal of the Linnean Society 151, pp. 1-339 : 17-18

HEPTAPACTES GEN. NOV.

Type species: Heptapactes quadrisaetosus sp. nov.

Diagnosis

Endoantennal conglobation. All coxal plates simple. Cephalothorax with a transverse groove not projected along the eye margin. Linea supraantennalis absent. Eyes composed of seven ommatidia. Second antenna flagellum two-jointed. Pleopod exopodites without distinct dorsolateral respiratory fields.

KEY TO THE AMERICAN GENERA OF ‘SCLEROPACTIDAE’ This key also includes some genera that have recently been classified as ‘Scleropactidae’ but that are excluded by the present phylogenetic analysis.

1. Cephalothorax with distinct median and lateral lobes parallel to the longitudinal axis. Coxal plates enlarged, directed outwards; species with clinger habitus...................................................... 2

1.* Cephalothorax with a frontal lamina vertical to the longitudinal axis; at most, small lateral lobes delimited. Coxal plates vertical, exoantennal or endoantennal conglobation ability...................................... 3

2. Cephalothorax with a deep, semicircular transverse furrow......................... Colomboscia pro parte

2.* Without such a furrow........................................................ Synuropus granulatus

3. Cephalothorax with distinct antennal lobes, second antenna apical cone short; flagellum two-jointed.................................................................................... Chileoniscus

3.* Cephalothorax without antennal lobes, second antenna apical cone long................................. 5

4. Eyes absent or represented only by a spot of dark pigment, without lenses. Pigment absent or present; transverse furrow behind the frontal shield always absent...................................................... 4

4.* Eyes present, with lenses; pigment always present; transverse furrow behind the frontal shield often present, may reach the inner margin of the eyes............................................................... 11

5. Exoantennal conglobation, frontal shield with two furrows for the second antennae.. Troglopactes botosaneanui

5.* Endoantennal conglobation...................................................................... 6

6. Coxal plate 1 with schisma at the posterior corner................................................... 9

6.* Coxal plate 1 with posterior corner simple.......................................................... 7

7. Flagellum of second antenna two-jointed; pigment absent; male pereiopod 7 ischium with ventrodistal process.................................................................................. Neosanfilippia

7.* Flagellum three-jointed......................................................................... 8

8. Visible portion of uropod sympodite about twice as long as broad; larger specimens with pigment........................................................................... Richardsoniscus portoricensis

8.* Visible portion of uropod sympodite about as long as broad; minute specimens 2–3 mm in length; without pigment.................................................................................. Colomboniscus

9. Second antennae stout, with fourth and fifth articles strongly enlarged, as long as broad, and with the shape of an equilateral triangle, one corner of which projects ventrally. Body without pigment, about 2 mm long............................................................................... Caecopactes minimus sp. nov.

9.* Second antennae not very stout, its articles longer than wide. Body larger, with or without pigment.> 5 mm ............................................................................................ 10

10. Uropod sympodite about as long as broad..................................... Sphaeroniscus gerstaeckeri

10.* Uropod sympodite about twice as long as broad.......................... Spherarmadillo Richardson, 1907

11. Coxal plates 2 and 3 with strongly concave lateral margin........................... Globopactes gen. nov.

11.* Coxal plates 2 and 3 with convex lateral margin................................................... 12

12. Coxal plate 1 with schisma..................................................................... 13

12.* Coxal plate 1 margin simple.................................................................... 14

13. Flagellum two-jointed................................................................... Circoniscus

13.* Flagellum three-jointed.............................................................. Sphaeroniscus

14. Flagellum two-jointed; male pereiopod 7 ischium with ventrodistal appendix................... Amazoniscus

14.* Flagellum three-jointed........................................................................ 15

15. Pleopod 1 and 2 exopodites with strongly wrinkled dorsal respiratory fields..................... Scleropactes

15.* Pleopod exopodites not distinctly wrinkled or without respiratory fields................................ 16

16. Transverse groove on cephalothorax in dorsal view evenly curved.................... Colomboscia pro parte

16.* Transverse groove medially straight and only laterally curved........................................ 17

17. Coxal plate 4 with acute angle.................................................................. 18

17,* Coxal plate 4 more rounded..................................................................... 19

18. Male pereiopod 7 merus with ventroproximal scaly tubercle.......................... Globopactes gen. nov.

18.* Male pereiopod 7 merus withiout such tubercle................................. Scleropactoides gen. nov.

19. Proximal article of maxilliped palp with only one large seta, male pereiopod 7 merus with triangular lobe on the frontal face; pleopod 1 endopodite distally smooth.................................... Colomboscia parva

19.* Proximal article of maxilliped palp with two large setae, male pereiopod 7 merus simple; pleopod 1 endopodite on the lateral face of the apex with small tubercles....................................... Scleropactes s.str.