Emphysemastix frampt Olsen & Enghoff, 2020

Olsen, Sissel Anna, Rosenmejer, Trine & Enghoff, Henrik, 2020, A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida), European Journal of Taxonomy 675, pp. 1-35 : 22-25

publication ID

https://doi.org/ 10.5852/ejt.2020.675

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persistent identifier


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scientific name

Emphysemastix frampt Olsen & Enghoff

sp. nov.

Emphysemastix frampt Olsen & Enghoff View in CoL sp. nov.


Figs 2 View Fig , 15B View Fig , 16–17 View Fig View Fig


Differs from the other species of the genus by having a rounded, rather than triangular paxillus. Nodal processes M and L both present.


The species is named after Kingseeker Frampt from the Dark Souls video game series, due to the gonopods’ resemblance to the creature. The name is to be treated as a noun in apposition.

Material examined


TANZANIA • ♂; Iringa Region, Mahenge District, West Kilombero Scarp FR, Nyumbanitu Mts, S of? Udekwa Village ; 07°48′ S, 36°21′ E; 1700 m a.s.l.; Dec. 1993; L.A. Hansen and J.O. Svendsen leg.; NHMD 621677 . GoogleMaps



SIZE AND SHAPE. A large member of the genus, rather wide compared to length. Body length ca 55 mm. Maximum width 13.1 mm. W/L ratio ca 24%.

COLOUR ( Fig. 16 View Fig ). Metazona orange-brown, with lighter colour on posterior part and on paranota; legs darker and more intensely/vividly orange. Prozona as metazona but without lighter areas.

BODY RINGS. Paranota set high on sides, dorsum moderately convex. Anterior edge of ring 2–4 straight, anterior edge of rings 5–20 with gradually larger posterior-facing edge. Torus present, but small and indistinct. Stricture poorly defined, but with sharp edge in front of anterior spiracles. Pleurosternal carinae present, knobby, distinct from ring 3, increasing in size towards ring 8–9, afterwards decreasing in size towards posterior end, indistinct on ring 18. Transverse sternal carinae as typical of the genus, but very small on anterior of ring 9. Anterior edge of paxillus rounded, with ridges.

LEGS. Relatively long ( Fig. 16 View Fig ).

HYPOPROCT. Paramediean tubercles small, not extended past edge of sclerite. Median projection large and elongated.

GONOPOD ( Figs 15B View Fig , 17 View Fig ). Gonocoxa with some paracannular setae, row of setae continuing on anterior side of distal end of gonocoxa. Prefemoral part with rounded ventral lobe. Telopodite endonodal. Nodal processes M and L similar in size; process L slightly curved mesad; process M slightly longer, distal end curved mesad. Postnodal telopodite relatively long, curved ventrad almost perpendicular to setose prefemoral region and directed first slightly mesad, then dorsad and expanding into subglobose, hollow enlargement, then curving mesad and forming a half-circle for the rest of its length. Subterminal process narrow and curved ventrad, solenomere curved dorsad, then ventrad.


Known only from the type locality in the Nyambanitu Mts, West Kilombero FR ( Fig. 2 View Fig ).


The gonopods of E. frampt sp. nov. are particularly similar to those of E. flavosignatus (compare Fig. 17 View Fig with Hoffman 2005: fig. 206). Processes M and L in E. frampt sp. nov. are, however, longer and more curved, and the nodus is larger, with more divergent sides in E. frampt sp. nov.

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