<document id="588414FF358B0C097099ADC0261315DF" ID-DOI="10.4202/app.2012.0076" ID-ISSN="1732-2421" ID-Zenodo-Dep="13277540" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_requiresApprovalFor="GgImagineBatch" checkinTime="1723148591768" checkinUser="felipe" docAuthor="Kustatscher, Evelyn, Dellantonio, Elio &amp; Van Konijnenburg-Van Cittert, Johanna H. A." docDate="2014" docId="D4702F298A7EFFECC3736175FF09D8AB" docLanguage="en" docName="ActaPalaeontolPol.59.3.741-755.pdf" docOrigin="Acta Palaeontologica Polonica 59 (3)" docSource="http://dx.doi.org/10.4202/app.2012.0076" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Cladophlebis undetermined" docType="treatment" docVersion="2" lastPageNumber="752" masterDocId="284957518A75FFE7C3116223FFE2DF0C" masterDocTitle="The ferns of the late Ladinian, Middle Triassic flora from Monte Agnello, Dolomites, Italy" masterLastPageNumber="755" masterPageNumber="741" pageNumber="752" updateTime="1723156092297" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DE4B3BFA4D36C2D5BEBE889DA200C532">The ferns of the late Ladinian, Middle Triassic flora from Monte Agnello, Dolomites, Italy</mods:title>
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<mods:namePart id="BF3BFC4571038752A21735DB85C6974E">Kustatscher, Evelyn</mods:namePart>
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<mods:namePart id="33C8541C544A949672AC04EE4A6F04B4">Dellantonio, Elio</mods:namePart>
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<mods:title id="1E67BBB5EBBB6F37F2278B2F590331E3">Acta Palaeontologica Polonica</mods:title>
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<paragraph id="5C669E3F8A7EFFECC3736175FEDADC7E" blockId="11.[98,312,854,882]" box="[98,312,854,882]" pageId="11" pageNumber="752">
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<taxonomicName id="9BD9E5BC8A7EFFECC3736175FEDADC7E" box="[98,312,854,882]" class="Polypodiopsida" family="Osmundaceae" genus="Cladophlebis" kingdom="Plantae" order="Osmundales" pageId="11" pageNumber="752" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="6EAD422D8A7EFFECC3736175FEEFDC7E" box="[98,269,854,882]" italics="true" pageId="11" pageNumber="752">Cladophlebis</emphasis>
sp.
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</heading>
</paragraph>
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<subSubSection id="14C3CDB48A7EFFECC37361A1FF09D8AB" pageId="11" pageNumber="752" type="description">
<paragraph id="5C669E3F8A7EFFECC37361A1FF29DC9B" blockId="11.[98,203,898,919]" box="[98,203,898,919]" pageId="11" pageNumber="752">
<figureCitation id="C4E282BA8A7EFFECC37361A1FF25DC9B" box="[98,199,898,919]" captionStart="Fig" captionStartId="10.[151,184,1409,1429]" captionTargetBox="[154,1533,162,1383]" captionTargetId="figure-234@10.[154,1533,162,1383]" captionTargetPageId="10" captionText="Fig. 6. Cladophlebis fronds from the tuff layer in the basal part of “explosion breccia” (late Ladinian, Middle Triassic) of Monte Agnello, Dolomites, N- Italy. A. Cladophlebis ladinica sp. nov., partially fused basal pinnules (MGP 194/6B). B–E. Cladophlebis sp. B. Bipinnate frond fragment (MGP 194/88). C. Pinnae with slightly falcate pinnules (MGP 194/87). D. Detail of the slightly imbricate pinnules (MGP 63/71). E. Possible basal pinnae with broad pinnules (MGP 194/89A). Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13277544" httpUri="https://zenodo.org/record/13277544/files/figure.png" pageId="11" pageNumber="752">Fig. 6B–E</figureCitation>
.
</paragraph>
<paragraph id="5C669E3F8A7EFFECC3736191FD61DCE7" blockId="11.[98,770,946,1003]" pageId="11" pageNumber="752">
<emphasis id="6EAD422D8A7EFFECC3736191FF01DCC7" box="[98,227,946,971]" italics="true" pageId="11" pageNumber="752">Material.—</emphasis>
MGP 63/71, 194/87, 194/88, 194/89A–B, Monte Agnello, Dolomites, N-Italy, early late Ladinian.
</paragraph>
<paragraph id="5C669E3F8A7EFFECC37361DEFE03DAF6" blockId="11.[98,771,1021,1530]" pageId="11" pageNumber="752">
<emphasis id="6EAD422D8A7EFFECC37361DEFEE7DB1A" box="[98,261,1021,1046]" italics="true" pageId="11" pageNumber="752">Description.—</emphasis>
The fronds are at least bipinnate. The fragments are up to 
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long and 
<quantity id="9B2133DA8A7EFFECC2DE663DFDFCDB3B" box="[463,542,1054,1079]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="11" pageNumber="752" unit="mm" value="55.0">55 mm</quantity>
wide (MGP 194/88; 
<figureCitation id="C4E282BA8A7EFFECC373661DFF59DB5B" box="[98,187,1086,1111]" captionStart="Fig" captionStartId="10.[151,184,1409,1429]" captionTargetBox="[154,1533,162,1383]" captionTargetId="figure-234@10.[154,1533,162,1383]" captionTargetPageId="10" captionText="Fig. 6. Cladophlebis fronds from the tuff layer in the basal part of “explosion breccia” (late Ladinian, Middle Triassic) of Monte Agnello, Dolomites, N- Italy. A. Cladophlebis ladinica sp. nov., partially fused basal pinnules (MGP 194/6B). B–E. Cladophlebis sp. B. Bipinnate frond fragment (MGP 194/88). C. Pinnae with slightly falcate pinnules (MGP 194/87). D. Detail of the slightly imbricate pinnules (MGP 63/71). E. Possible basal pinnae with broad pinnules (MGP 194/89A). Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13277544" httpUri="https://zenodo.org/record/13277544/files/figure.png" pageId="11" pageNumber="752">Fig. 6B</figureCitation>
). The primary rachis is 
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thick. Pinnae arise suboppositely and at an acute angle (45–50°) from the rachis, but then curve slightly outwards. Pinnae fragments are up to 
<quantity id="9B2133DA8A7EFFECC37366BDFF53DBBB" box="[98,177,1182,1207]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="11" pageNumber="752" unit="mm" value="55.0">55 mm</quantity>
long and generally 
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wide; just in one case the pinnae fragments are broader (MGP 194/89A; 
<figureCitation id="C4E282BA8A7EFFECC19A669CFD3DDBD4" box="[651,735,1215,1240]" captionStart="Fig" captionStartId="10.[151,184,1409,1429]" captionTargetBox="[154,1533,162,1383]" captionTargetId="figure-234@10.[154,1533,162,1383]" captionTargetPageId="10" captionText="Fig. 6. Cladophlebis fronds from the tuff layer in the basal part of “explosion breccia” (late Ladinian, Middle Triassic) of Monte Agnello, Dolomites, N- Italy. A. Cladophlebis ladinica sp. nov., partially fused basal pinnules (MGP 194/6B). B–E. Cladophlebis sp. B. Bipinnate frond fragment (MGP 194/88). C. Pinnae with slightly falcate pinnules (MGP 194/87). D. Detail of the slightly imbricate pinnules (MGP 63/71). E. Possible basal pinnae with broad pinnules (MGP 194/89A). Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13277544" httpUri="https://zenodo.org/record/13277544/files/figure.png" pageId="11" pageNumber="752">Fig. 6E</figureCitation>
). It could belong to a basal frond fragment. The rachis of the pinnae is 
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wide. The pinnules are lanceolate to slightly falcate with a rounded apex and a broad base. The pinnules are densely inserted alternately to suboppositely at an angle of 70–80°. The pinnules are 
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long and 
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wide (MGP 194/87; 
<figureCitation id="C4E282BA8A7EFFECC24567A3FE52DA95" box="[340,432,1408,1433]" captionStart="Fig" captionStartId="10.[151,184,1409,1429]" captionTargetBox="[154,1533,162,1383]" captionTargetId="figure-234@10.[154,1533,162,1383]" captionTargetPageId="10" captionText="Fig. 6. Cladophlebis fronds from the tuff layer in the basal part of “explosion breccia” (late Ladinian, Middle Triassic) of Monte Agnello, Dolomites, N- Italy. A. Cladophlebis ladinica sp. nov., partially fused basal pinnules (MGP 194/6B). B–E. Cladophlebis sp. B. Bipinnate frond fragment (MGP 194/88). C. Pinnae with slightly falcate pinnules (MGP 194/87). D. Detail of the slightly imbricate pinnules (MGP 63/71). E. Possible basal pinnae with broad pinnules (MGP 194/89A). Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13277544" httpUri="https://zenodo.org/record/13277544/files/figure.png" pageId="11" pageNumber="752">Fig. 6C</figureCitation>
). The midrib is distinct. The secondary veins arise at an acute angle (c. 50°), bifurcate once halfway along the lamina and perhaps once again near the margin (MGP 63/71; 
<figureCitation id="C4E282BA8A7EFFECC26A67C2FE36DAF6" box="[379,468,1505,1530]" captionStart="Fig" captionStartId="10.[151,184,1409,1429]" captionTargetBox="[154,1533,162,1383]" captionTargetId="figure-234@10.[154,1533,162,1383]" captionTargetPageId="10" captionText="Fig. 6. Cladophlebis fronds from the tuff layer in the basal part of “explosion breccia” (late Ladinian, Middle Triassic) of Monte Agnello, Dolomites, N- Italy. A. Cladophlebis ladinica sp. nov., partially fused basal pinnules (MGP 194/6B). B–E. Cladophlebis sp. B. Bipinnate frond fragment (MGP 194/88). C. Pinnae with slightly falcate pinnules (MGP 194/87). D. Detail of the slightly imbricate pinnules (MGP 63/71). E. Possible basal pinnae with broad pinnules (MGP 194/89A). Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13277544" httpUri="https://zenodo.org/record/13277544/files/figure.png" pageId="11" pageNumber="752">Fig. 6D</figureCitation>
).
</paragraph>
<paragraph id="5C669E3F8A7EFFECC373642FFF09D8AB" blockId="11.[98,771,1548,1959]" pageId="11" pageNumber="752">
<emphasis id="6EAD422D8A7EFFECC373642FFF07D929" box="[98,229,1548,1573]" italics="true" pageId="11" pageNumber="752">Remarks.—</emphasis>
These remains are very fragmentary and too badly preserved to be assigned to a new or established species. They resemble the Late Triassic species 
<taxonomicName id="9BD9E5BC8A7EFFECC1A2646FFE36D989" authority="(Presl, 1838) Van" authorityName="Van" baseAuthorityName="Presl" baseAuthorityYear="1838" class="Polypodiopsida" family="Osmundaceae" genus="Cladophlebis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Osmundales" pageId="11" pageNumber="752" phylum="Tracheophyta" rank="species" species="remota">
<emphasis id="6EAD422D8A7EFFECC1A2646FFEE8D989" italics="true" pageId="11" pageNumber="752">Cladophlebis remota</emphasis>
(Presl, 1838) Van
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Konijnenburg-van Cittert, Van Konijnenburg-van Cittert, Kustatscher, and Wachtler, 2006, but in the latter species the pinnules are more closely inserted and the apex is acute. 
<taxonomicName id="9BD9E5BC8A7EFFECC2FE64EEFECCD80A" authority="Brongniart, 1828" authorityName="Brongniart" authorityYear="1828" class="Polypodiopsida" family="Osmundaceae" genus="Cladophlebis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Osmundales" pageId="11" pageNumber="752" phylum="Tracheophyta" rank="species" species="nebbensis">
<emphasis id="6EAD422D8A7EFFECC2FE64EEFCE1D9EA" box="[495,771,1741,1766]" italics="true" pageId="11" pageNumber="752">Cladophlebis nebbensis</emphasis>
Brongniart, 1828
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) Nathorst, 1876, a typical Rhaeto-Liassic species from Europe and Asia is similar in its once forked secondary veins and slightly restricted pinnule bases, but the pinnules dimensions in 
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<emphasis id="6EAD422D8A7EFFECC28D656DFDCCD86B" box="[412,558,1870,1895]" italics="true" pageId="11" pageNumber="752">C. nebbensis</emphasis>
</taxonomicName>
are larger (around 
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long) and the rachis is relatively thin (
<bibRefCitation id="3848E3CE8A7EFFECC190654DFF3CD8AB" author="Schweitzer, H. - J. &amp; Van Konijnenburg-van &amp; Cittert, J. H. A. &amp; Van der Burgh, J." pageId="11" pageNumber="752" pagination="103 - 192" refId="ref11899" refString="Schweitzer, H. - J., Van Konijnenburg-van Cittert, J. H. A., and Van der Burgh, J. 1997. The Rhaeto-Liassic flora of Iran and Afghanistan. 10. Bryophyta, Lycophyta, Sphenophyta, Pterophyta - Eusporangiatae and Protosporangiatae. Palaeontographica B 243: 103 - 192." type="journal article" year="1997">Schweitzer et al. 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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