A revision and phylogenetic analysis of the spider genus Coptoprepes Simon (Araneae: Anyphaenidae, Amaurobioidinae)
Author
Werenkraut, Victoria
Author
Ramírez, Martín J.
text
Zootaxa
2009
2212
1
40
journal article
40616
10.5281/zenodo.275145
344a109d-a561-4054-97b7-e1850acc321f
1175-5326
275145
Coptoprepes ecotono
new species
Figs. 5–7
,
22
a, 23a–c, 24a–c
Type
material:
Holotype
:
male from
Argentina
, Río Negro province, Bariloche, ca. 41º
0 0 S
,
71º30’ W
,
March 1947
, A. G. Giai (MACN-Ar 2218; temporary preparations MJR-01341, VIW-00014).
Paratypes
:
female from
Argentina
, Neuquén province,
Nahuel
Huapi Natl. Park, Lago Espejo, ca.
40º 30’ S
,
71º 30’ W
,
21 January 1985
, M. J. Ramírez (MACN-Ar 20320; temporary preparations MJR-01342, VIW-00013).
Etymology:
This new species is dedicated to V. Werenkraut’s colleagues and friends from Laboratorio
Ecotono
, Bariloche,
Argentina
.
Diagnosis:
Females are very similar to those of
C. valdiviensis
by the rather flat epigyne with rectangular, unsclerotized median field (
Fig.
7
g), but can be distinguished by having a longitudinally oriented profile of the anterior portion of the copulatory ducts (
Fig. 24
a–c). Males are similar to those of
C. campanensis
n. sp.
by (virtually) lacking a RTA, but have a much smaller, not bifid median apophysis (
Figs. 7
a–f, 23a–c).
FIGURE 4.
Coptoprepes valdiviensis
Ramírez.
a
female dorsal habitus;
b
epigyne, ventral view;
c
male left palp, ventral view;
d
male left palp, retrolateral view. Preparations, female MJR-01347, male MJR-01328.
FIGURE 5.
Coptoprepes ecotono
new species
.
a–d
male holotype (
a
dorsal habitus;
b
prosoma, dorsal view;
c
ventral habitus, ventral view;
d
prosoma, ventral view).
FIGURE 6.
Coptoprepes ecotono
new species
.
a–d
female paratype (
a
dorsal habitus;
b
prosoma, dorsal view;
c
ventral habitus, ventral view;
d
prosoma, ventral view).
FIGURE 7.
Coptoprepes ecotono
new species
.
a–f
male,
g–h
female paratype (
a
left palp, male holotype, prolateral view,
b
same, ventral view,
c
same, retrolateral view;
d
copulatory bulb, preparation MJR-01343, prolateral view,
e
same, apical view,
f
same, retrolateral view;
g
epigyne, ventral view;
h
vulva).
Description:
Male
(
holotype
): Prosoma length 1.86, width 1.36, narrower in front than in female. Tibia/ metatarsus length: I, 1.06/0.91; II, 0.92/0.84; III, 0.64/0.68; IV, 1.17/1.27. Chelicerae slightly smaller than those of female, with four teeth on retromargin, three on promargin (promargin observed from MACN-Ar 2227). Sternum length 1.05, width 0.78. Spines:
I
, femur d 1-1-1, p 2ap; tibia v 0-2-p1; metatarsus
v 2
bas.
II
, femur d 1-1-1, p d1ap; tibia v 0-r1-p1; metatarsus
v 2
bas.
III
, femur d 1-1-1, p and r 0-d1-d1; patella r d1; tibia v p1-2-2, p 1-d1-1-0, r d1-1, d 0; metatarsus
v 2
-p1-2 plus apical comb, p and r d1-1-1, d 0-p1-
2
.
IV
, femur d 1-1-1, p 0-d1-d1, r d1ap; patella r d1; tibia
v 2-2
-2, p and r 1-d1-1-0, d 0; metatarsus
v 2-2
-2 plus apical comb, p and r d1-1-1, d 0-2-2. Opisthosoma length 2.20, spiracle–epigastrium 1.15, spiracle–spinnerets 0.18. Color in ethanol (
Fig. 5
): As in female, but paler, specimen faded (as well as male MACN-Ar 2227). Palp (
Fig. 7
): Tibia short, width/length 1.00, RTA reduced to short acute relict. Cymbial conductor wide. Tegulum only visible in retrolateral view. Sperm duct with conspicuous loop on anterior-dorsal margin. Embolus long, thin, basal process elongate, flat, transparent. Median apophysis thin, hook-shaped. Primary conductor with long canal, ventral border tiny, cuticular hair-like extensions, tip acute, elongate, directed forward. Secondary conductor well sclerotized, apical, without canal, contiguous to anterior-dorsal sperm duct loop. Paramedian apophysis absent, area only slightly sclerotized (
Fig. 23
a–c).
Female
(
paratype
): Prosoma length 1.82, width 1.29. Tibia/metatarsus length: I, 0.93/0.80; II, 0.83/0.78; III, 0.72/0.86; IV, 1.11/131. Chelicerae with four teeth on retromargin, four on promargin. Sternum length 1.05, width 0.76. Spines: leg
I
, femur d 1-1-1, p 2ap; tibia
v 2-2
-2; metatarsus
v 2
bas.
II
, femur d 1-1-1, p d1ap; tibia v r1-r1-p1; metatarsus
v 2
bas.
III
, femur d 1-1-1, p and r 0-d1-d1; patella r d1; tibia v p1-p1-2, p 1- d1-1-0, r d1-1, d r1bas; metatarsus
v 2
-0-p1 plus apical comb, p and r d1-1-1, d 0-p1-
2
.
IV
, femur d 1-1-1, p and r d1ap; patella r d1; tibia v p1-2-2, p and r 1-d1-1-0, d r1bas; metatarsus
v 2-2
-p1 plus apical comb, p and r d1-1-1, d 0-p1-2. Color in ethanol (
Fig. 6
): Prosoma brown, legs brown, darker distally. Opisthosoma pale brown, dorsal pattern brown, including posterior chevrons. Opisthosoma length 2.55, epigastrium–spiracle 1.19, spiracle–spinnerets 0.15. Epigyne: Lateral lobes separate, median field not sclerotized (
Fig.
7
g). Copulatory openings very close to epigastric furrow. Copulatory ducts slightly wrapped. Ducts of accessory bulbs short. Spermathecae with convoluted chambers (
Figs. 7
h, 24a–c).
Variability:
Spines in male: I, tibia v 0-p1-p
1. II
, tibia v 0-2-p
1. III
, metatarsus
v 2
-r1-2 or 2-2-2.
Distribution:
Known from Neuquén and Río Negro provinces in
Argentina
.
Other material examined:
ARGENTINA
:
same locality as
holotype
,
1 male
(MACN-Ar 2227; temporary preparation MJR-01343); same data as
paratype
[1 immature] (MACN-Ar).