A revision and phylogenetic analysis of the spider genus Coptoprepes Simon (Araneae: Anyphaenidae, Amaurobioidinae) Author Werenkraut, Victoria Author Ramírez, Martín J. text Zootaxa 2009 2212 1 40 journal article 40616 10.5281/zenodo.275145 344a109d-a561-4054-97b7-e1850acc321f 1175-5326 275145 Coptoprepes ecotono new species Figs. 5–7 , 22 a, 23a–c, 24a–c Type material: Holotype : male from Argentina , Río Negro province, Bariloche, ca. 41º 0 0 S , 71º30’ W , March 1947 , A. G. Giai (MACN-Ar 2218; temporary preparations MJR-01341, VIW-00014). Paratypes : female from Argentina , Neuquén province, Nahuel Huapi Natl. Park, Lago Espejo, ca. 40º 30’ S , 71º 30’ W , 21 January 1985 , M. J. Ramírez (MACN-Ar 20320; temporary preparations MJR-01342, VIW-00013). Etymology: This new species is dedicated to V. Werenkraut’s colleagues and friends from Laboratorio Ecotono , Bariloche, Argentina . Diagnosis: Females are very similar to those of C. valdiviensis by the rather flat epigyne with rectangular, unsclerotized median field ( Fig. 7 g), but can be distinguished by having a longitudinally oriented profile of the anterior portion of the copulatory ducts ( Fig. 24 a–c). Males are similar to those of C. campanensis n. sp. by (virtually) lacking a RTA, but have a much smaller, not bifid median apophysis ( Figs. 7 a–f, 23a–c). FIGURE 4. Coptoprepes valdiviensis Ramírez. a female dorsal habitus; b epigyne, ventral view; c male left palp, ventral view; d male left palp, retrolateral view. Preparations, female MJR-01347, male MJR-01328. FIGURE 5. Coptoprepes ecotono new species . a–d male holotype ( a dorsal habitus; b prosoma, dorsal view; c ventral habitus, ventral view; d prosoma, ventral view). FIGURE 6. Coptoprepes ecotono new species . a–d female paratype ( a dorsal habitus; b prosoma, dorsal view; c ventral habitus, ventral view; d prosoma, ventral view). FIGURE 7. Coptoprepes ecotono new species . a–f male, g–h female paratype ( a left palp, male holotype, prolateral view, b same, ventral view, c same, retrolateral view; d copulatory bulb, preparation MJR-01343, prolateral view, e same, apical view, f same, retrolateral view; g epigyne, ventral view; h vulva). Description: Male ( holotype ): Prosoma length 1.86, width 1.36, narrower in front than in female. Tibia/ metatarsus length: I, 1.06/0.91; II, 0.92/0.84; III, 0.64/0.68; IV, 1.17/1.27. Chelicerae slightly smaller than those of female, with four teeth on retromargin, three on promargin (promargin observed from MACN-Ar 2227). Sternum length 1.05, width 0.78. Spines: I , femur d 1-1-1, p 2ap; tibia v 0-2-p1; metatarsus v 2 bas. II , femur d 1-1-1, p d1ap; tibia v 0-r1-p1; metatarsus v 2 bas. III , femur d 1-1-1, p and r 0-d1-d1; patella r d1; tibia v p1-2-2, p 1-d1-1-0, r d1-1, d 0; metatarsus v 2 -p1-2 plus apical comb, p and r d1-1-1, d 0-p1- 2 . IV , femur d 1-1-1, p 0-d1-d1, r d1ap; patella r d1; tibia v 2-2 -2, p and r 1-d1-1-0, d 0; metatarsus v 2-2 -2 plus apical comb, p and r d1-1-1, d 0-2-2. Opisthosoma length 2.20, spiracle–epigastrium 1.15, spiracle–spinnerets 0.18. Color in ethanol ( Fig. 5 ): As in female, but paler, specimen faded (as well as male MACN-Ar 2227). Palp ( Fig. 7 ): Tibia short, width/length 1.00, RTA reduced to short acute relict. Cymbial conductor wide. Tegulum only visible in retrolateral view. Sperm duct with conspicuous loop on anterior-dorsal margin. Embolus long, thin, basal process elongate, flat, transparent. Median apophysis thin, hook-shaped. Primary conductor with long canal, ventral border tiny, cuticular hair-like extensions, tip acute, elongate, directed forward. Secondary conductor well sclerotized, apical, without canal, contiguous to anterior-dorsal sperm duct loop. Paramedian apophysis absent, area only slightly sclerotized ( Fig. 23 a–c). Female ( paratype ): Prosoma length 1.82, width 1.29. Tibia/metatarsus length: I, 0.93/0.80; II, 0.83/0.78; III, 0.72/0.86; IV, 1.11/131. Chelicerae with four teeth on retromargin, four on promargin. Sternum length 1.05, width 0.76. Spines: leg I , femur d 1-1-1, p 2ap; tibia v 2-2 -2; metatarsus v 2 bas. II , femur d 1-1-1, p d1ap; tibia v r1-r1-p1; metatarsus v 2 bas. III , femur d 1-1-1, p and r 0-d1-d1; patella r d1; tibia v p1-p1-2, p 1- d1-1-0, r d1-1, d r1bas; metatarsus v 2 -0-p1 plus apical comb, p and r d1-1-1, d 0-p1- 2 . IV , femur d 1-1-1, p and r d1ap; patella r d1; tibia v p1-2-2, p and r 1-d1-1-0, d r1bas; metatarsus v 2-2 -p1 plus apical comb, p and r d1-1-1, d 0-p1-2. Color in ethanol ( Fig. 6 ): Prosoma brown, legs brown, darker distally. Opisthosoma pale brown, dorsal pattern brown, including posterior chevrons. Opisthosoma length 2.55, epigastrium–spiracle 1.19, spiracle–spinnerets 0.15. Epigyne: Lateral lobes separate, median field not sclerotized ( Fig. 7 g). Copulatory openings very close to epigastric furrow. Copulatory ducts slightly wrapped. Ducts of accessory bulbs short. Spermathecae with convoluted chambers ( Figs. 7 h, 24a–c). Variability: Spines in male: I, tibia v 0-p1-p 1. II , tibia v 0-2-p 1. III , metatarsus v 2 -r1-2 or 2-2-2. Distribution: Known from Neuquén and Río Negro provinces in Argentina . Other material examined: ARGENTINA : same locality as holotype , 1 male (MACN-Ar 2227; temporary preparation MJR-01343); same data as paratype [1 immature] (MACN-Ar).