<document id="64D4C9419BBD8B38E55EBC441D42B629" ID-DOI="10.5281/zenodo.5325891" ID-ISSN="0253-116X" ID-Zenodo-Dep="5325891" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630152202360" checkinUser="carolina" docAuthor="Assing, V." docDate="2011" docId="AC30A044BC4BFFB0FF01FCAFFC7D204C" docLanguage="en" docName="LinzerbiolBeitr.2011.43.2.1135-1158.pdf" docOrigin="Linzer biologische Beiträge 43 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Geostiba (Tropogastrosipalia) burdurica Assing 2011, nov.sp." docType="treatment" docVersion="4" lastPageNumber="1147" masterDocId="5009D83CBC40FFBCFFB3FFA2FFD0260A" masterDocTitle="A revision of Geostiba of the West Palaearctic region. XXI. Eight new species from Turkey and the Caucasus, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="1158" masterPageNumber="1135" pageNumber="1146" updateTime="1699165521890" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EC40D00776BB2C073728B598287DB567">A revision of Geostiba of the West Palaearctic region. XXI. Eight new species from Turkey and the Caucasus, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="29708E89F3CF101FA16200D607C4231A">Assing, V.</mods:namePart>
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<subSubSection id="6C8342D9BC4BFFB7FF01FCAFFF3C2599" pageId="11" pageNumber="1146" type="nomenclature">
<paragraph id="24261152BC4BFFB7FF01FCAFFC84252F" blockId="11.[178,852,781,806]" box="[178,852,781,806]" pageId="11" pageNumber="1146">
<heading id="7F6EA63EBC4BFFB7FF01FCAFFC84252F" bold="true" box="[178,852,781,806]" fontSize="9" level="2" pageId="11" pageNumber="1146" reason="3">
<emphasis id="16EDCD40BC4BFFB7FF01FCAFFD17252C" bold="true" box="[178,711,781,806]" pageId="11" pageNumber="1146">
<taxonomicName id="E3996AD1BC4BFFB7FF01FCAFFDA0252F" ID-CoL="6KCVT" authority="Assing, 2011" authorityName="Assing" authorityYear="2011" box="[178,624,781,805]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1146" phylum="Arthropoda" rank="species" species="burdurica" status="sp. nov." subGenus="Tropogastrosipalia">
<emphasis id="16EDCD40BC4BFFB7FF01FCAFFEC3252F" bold="true" box="[178,275,781,805]" italics="true" pageId="11" pageNumber="1146">Geostiba</emphasis>
(
<emphasis id="16EDCD40BC4BFFB7FE91FCAFFE22252F" bold="true" box="[290,498,781,805]" italics="true" pageId="11" pageNumber="1146">Tropogastrosipalia</emphasis>
) 
<emphasis id="16EDCD40BC4BFFB7FDB1FCAFFDA0252F" bold="true" box="[514,624,781,805]" italics="true" pageId="11" pageNumber="1146">burdurica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DDE703BBC4BFFB7FDC4FCACFD17252C" box="[631,711,782,806]" pageId="11" pageNumber="1146" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="BCA20DD7BC4BFFB7FD65FCACFC9C252F" box="[726,844,782,805]" captionStart="Figs 26-32" captionStartId="12.[178,218,1305,1327]" captionTargetBox="[178,1076,433,1281]" captionTargetId="figure-278@12.[178,1085,433,1297]" captionTargetPageId="12" captionText="Figs 26-32: Geostiba burdurica nov.sp. (26, 29: holotype): (26) male forebody; (27) male abdominal apex in lateral view; (28) posterior portion of male abdominal tergite VII in antero-dorsal view; (29-30) median lobe of aedeagus in lateral view; (31-32) spermatheca. Scale bars: 26: 0.5 mm; 27- 32: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5325913" httpUri="https://zenodo.org/record/5325913/files/figure.png" pageId="11" pageNumber="1146">Figs 26-32</figureCitation>
)
</heading>
</paragraph>
<paragraph id="24261152BC4BFFB7FF01FC99FF3C2599" blockId="11.[178,1085,827,1353]" pageId="11" pageNumber="1146">
T y p e m a t e r i a l: 
<typeStatus id="FB22AFF0BC4BFFB7FE18FC99FDD02545" box="[427,512,827,847]" pageId="11" pageNumber="1146" type="holotype">Holotype</typeStatus>
: &quot;TR [8], 
<collectingRegion id="E65DDFB0BC4BFFB7FD39FC99FD1D2545" box="[650,717,827,847]" country="Turkey" name="Burdur" pageId="11" pageNumber="1146">Burdur</collectingRegion>
, W Çeltikçi Geç., 
<quantity id="E361BCB7BC4BFFB7FC3BFC99FC172545" box="[904,967,827,847]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="1146" unit="km" value="21.0">21 km</quantity>
SE 
<collectingRegion id="E65DDFB0BC4BFFB7FC44FC99FBEA2545" box="[1015,1082,827,847]" country="Turkey" name="Burdur" pageId="11" pageNumber="1146">Burdur</collectingRegion>
, 
<geoCoordinate id="41AD7795BC4BFFB7FF01FCF0FECE256C" box="[178,286,850,870]" degrees="37" direction="north" minutes="33" orientation="latitude" pageId="11" pageNumber="1146" precision="15" seconds="37" value="37.560276">37°33'37''N</geoCoordinate>
, 
<geoCoordinate id="41AD7795BC4BFFB7FE9AFCF0FE42256C" box="[297,402,850,870]" degrees="30" direction="east" minutes="23" orientation="longitude" pageId="11" pageNumber="1146" precision="15" seconds="49" value="30.396944">30°23'49''E</geoCoordinate>
, 
<quantity id="E361BCB7BC4BFFB7FE2DFCF0FE37256C" box="[414,487,850,870]" metricMagnitude="3" metricUnit="m" metricValue="1.419" pageId="11" pageNumber="1146" unit="m" value="1419.0">1419 m</quantity>
, Zypr., Eichen, 
<date id="50273792BC4BFFB7FDCEFCF0FD0D256C" box="[637,733,850,870]" pageId="11" pageNumber="1146" value="2011-02-16">16.II.2011</date>
, M. Schülke / 
<typeStatus id="FB22AFF0BC4BFFB7FCD9FCF0FC19256C" box="[874,969,850,870]" pageId="11" pageNumber="1146" type="holotype">Holotypus</typeStatus>
<taxonomicName id="E3996AD1BC4BFFB7FC58FCF2FEDD2576" authority="Assing, 2011" authorityName="Assing" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1146" phylum="Arthropoda" rank="species" species="burdurica" status="sp. nov.">
<emphasis id="16EDCD40BC4BFFB7FC58FCF2FEDD2576" italics="true" pageId="11" pageNumber="1146">Geostiba burdurica</emphasis>
</taxonomicName>
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det. V. Assing 2011&quot; (cAss). 
<typeStatus id="FB22AFF0BC4BFFB7FDE6FCCBFD612577" box="[597,689,873,893]" pageId="11" pageNumber="1146" type="paratype">Paratypes</typeStatus>
: 6, 5: same data as 
<typeStatus id="FB22AFF0BC4BFFB7FC1DFCCBFC2D2577" box="[942,1021,873,893]" pageId="11" pageNumber="1146" type="holotype">holotype</typeStatus>
(cSch, cAss).
</paragraph>
</subSubSection>
<subSubSection id="6C8342D9BC4BFFB0FF01FC38FC7D204C" lastPageId="12" lastPageNumber="1147" pageId="11" pageNumber="1146" type="description">
<paragraph id="24261152BC4BFFB7FF01FC38FC15220D" blockId="11.[178,1085,827,1353]" pageId="11" pageNumber="1146">
D e s c r i p t i o n: Body length 
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. Coloration: head dark-brown to blackish; pronotum and elytra reddish, distinctly contrasting with the head; abdomen reddishbrown, with segments VI-VII or V-VII more or less extensively infuscate; legs yellowish; antennae reddish to brown, with the basal antennomeres pale-reddish.
</paragraph>
<paragraph id="24261152BC4BFFB7FF01FBADFC9A2255" blockId="11.[178,1085,827,1353]" pageId="11" pageNumber="1146">
Head approximately as wide as long; punctation sparse and extremely fine, barely noticeable; interstices with shallow microsculpture; eyes weakly convex and approximately half as long as postocular portion in dorsal view (
<figureCitation id="BCA20DD7BC4BFFB7FD41FBEBFCEE2255" box="[754,830,1097,1119]" captionStart="Figs 26-32" captionStartId="12.[178,218,1305,1327]" captionTargetBox="[178,1076,433,1281]" captionTargetId="figure-278@12.[178,1085,433,1297]" captionTargetPageId="12" captionText="Figs 26-32: Geostiba burdurica nov.sp. (26, 29: holotype): (26) male forebody; (27) male abdominal apex in lateral view; (28) posterior portion of male abdominal tergite VII in antero-dorsal view; (29-30) median lobe of aedeagus in lateral view; (31-32) spermatheca. Scale bars: 26: 0.5 mm; 27- 32: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5325913" httpUri="https://zenodo.org/record/5325913/files/figure.png" pageId="11" pageNumber="1146">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="24261152BC4BFFB7FF01FBC5FCFE2290" blockId="11.[178,1085,827,1353]" pageId="11" pageNumber="1146">Pronotum with weakly pronounced sexual dimorphism; punctation very fine and barely noticeable; microsculpture more pronounced than that of head.</paragraph>
<paragraph id="24261152BC4BFFB7FF01FB03FECB22DE" blockId="11.[178,1085,827,1353]" pageId="11" pageNumber="1146">Elytra with moderately pronounced sexual dimorphism, approximately half as long as pronotum.</paragraph>
<paragraph id="24261152BC4BFFB7FF01FB7EFBE12343" blockId="11.[178,1085,827,1353]" pageId="11" pageNumber="1146">Abdomen slightly broader than elytra; tergites III-IV and VII with sexual dimorphism; punctation fine to moderately fine, moderately dense on anterior tergites and very sparse on posterior tergites; interstices with distinct, but shallow microsculpture; posterior margin of tergite VII usually with, rarely without very narrow rudiment of a palisade fringe.</paragraph>
<paragraph id="24261152BC4BFFB0FF61FAFAFE1E2701" blockId="11.[178,1085,1368,1586]" lastBlockId="12.[178,1085,217,267]" lastPageId="12" lastPageNumber="1147" pageId="11" pageNumber="1146">
(with fully developed secondary sexual characters): pronotum weakly oblong, approximately 1.05 times as long as broad, posterior margin strongly convex in the middle; elytra slightly less than half as long as pronotum, very shallowly impressed at most, with moderately granulose punctation, and with moderately pronounced sutural carinae extending along anterior 2/3 of suture (
<figureCitation id="BCA20DD7BC4BFFB7FD9CFA6AFDAD23D4" box="[559,637,1480,1502]" captionStart="Figs 26-32" captionStartId="12.[178,218,1305,1327]" captionTargetBox="[178,1076,433,1281]" captionTargetId="figure-278@12.[178,1085,433,1297]" captionTargetPageId="12" captionText="Figs 26-32: Geostiba burdurica nov.sp. (26, 29: holotype): (26) male forebody; (27) male abdominal apex in lateral view; (28) posterior portion of male abdominal tergite VII in antero-dorsal view; (29-30) median lobe of aedeagus in lateral view; (31-32) spermatheca. Scale bars: 26: 0.5 mm; 27- 32: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5325913" httpUri="https://zenodo.org/record/5325913/files/figure.png" pageId="11" pageNumber="1146">Fig. 26</figureCitation>
); abdominal tergites II and III with median tubercles, both of them situated close to the anterior tergal impressions; process of tergite VII broadly triangular in lateral view (
<figureCitation id="BCA20DD7BC4BFFB7FDE5F9A2FD77201C" box="[598,679,1536,1558]" captionStart="Figs 26-32" captionStartId="12.[178,218,1305,1327]" captionTargetBox="[178,1076,433,1281]" captionTargetId="figure-278@12.[178,1085,433,1297]" captionTargetPageId="12" captionText="Figs 26-32: Geostiba burdurica nov.sp. (26, 29: holotype): (26) male forebody; (27) male abdominal apex in lateral view; (28) posterior portion of male abdominal tergite VII in antero-dorsal view; (29-30) median lobe of aedeagus in lateral view; (31-32) spermatheca. Scale bars: 26: 0.5 mm; 27- 32: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5325913" httpUri="https://zenodo.org/record/5325913/files/figure.png" pageId="11" pageNumber="1146">Fig. 27</figureCitation>
), parallel, convex (cross-section), and apically rounded in antero-dorsal view (
<figureCitation id="BCA20DD7BC4BFFB7FDFBF9BEFD442038" box="[584,660,1564,1586]" captionStart="Figs 26-32" captionStartId="12.[178,218,1305,1327]" captionTargetBox="[178,1076,433,1281]" captionTargetId="figure-278@12.[178,1085,433,1297]" captionTargetPageId="12" captionText="Figs 26-32: Geostiba burdurica nov.sp. (26, 29: holotype): (26) male forebody; (27) male abdominal apex in lateral view; (28) posterior portion of male abdominal tergite VII in antero-dorsal view; (29-30) median lobe of aedeagus in lateral view; (31-32) spermatheca. Scale bars: 26: 0.5 mm; 27- 32: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5325913" httpUri="https://zenodo.org/record/5325913/files/figure.png" pageId="11" pageNumber="1146">Fig. 28</figureCitation>
); median lobe of aedeagus approximately 
<quantity id="E361BCB7BC4CFFB0FF01FF7BFEDC26E5" box="[178,268,217,239]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="1147" unit="mm" value="0.28">0.28 mm</quantity>
long, cristal process narrow and obliquely projecting away from ventral process in lateral view (
<figureCitation id="BCA20DD7BC4CFFB0FEE3FF57FE132701" box="[336,451,245,267]" captionStart="Figs 26-32" captionStartId="12.[178,218,1305,1327]" captionTargetBox="[178,1076,433,1281]" captionTargetId="figure-278@12.[178,1085,433,1297]" captionTargetPageId="12" captionText="Figs 26-32: Geostiba burdurica nov.sp. (26, 29: holotype): (26) male forebody; (27) male abdominal apex in lateral view; (28) posterior portion of male abdominal tergite VII in antero-dorsal view; (29-30) median lobe of aedeagus in lateral view; (31-32) spermatheca. Scale bars: 26: 0.5 mm; 27- 32: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5325913" httpUri="https://zenodo.org/record/5325913/files/figure.png" pageId="12" pageNumber="1147">Figs 29-30</figureCitation>
).
</paragraph>
<paragraph id="24261152BC4CFFB0FF74FEB8FD9F2741" blockId="12.[178,1085,281,331]" pageId="12" pageNumber="1147">
: pronotum weakly transverse; elytra unmodified and slightly more than half as long as pronotum; spermatheca as in 
<figureCitation id="BCA20DD7BC4CFFB0FE68FE97FD9F2741" box="[475,591,309,331]" captionStart="Figs 26-32" captionStartId="12.[178,218,1305,1327]" captionTargetBox="[178,1076,433,1281]" captionTargetId="figure-278@12.[178,1085,433,1297]" captionTargetPageId="12" captionText="Figs 26-32: Geostiba burdurica nov.sp. (26, 29: holotype): (26) male forebody; (27) male abdominal apex in lateral view; (28) posterior portion of male abdominal tergite VII in antero-dorsal view; (29-30) median lobe of aedeagus in lateral view; (31-32) spermatheca. Scale bars: 26: 0.5 mm; 27- 32: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5325913" httpUri="https://zenodo.org/record/5325913/files/figure.png" pageId="12" pageNumber="1147">Figs 31-32.</figureCitation>
</paragraph>
<paragraph id="24261152BC4CFFB0FF01FEF8FC8B27AD" blockId="12.[178,1085,345,423]" pageId="12" pageNumber="1147">
E t y m o l o g y: The specific epithet is a latinized adjective derived from the name of the province where the 
<typeStatus id="FB22AFF0BC4CFFB0FE10FED4FE1F2781" box="[419,463,374,395]" pageId="12" pageNumber="1147">type</typeStatus>
locality is situated. It refers to the fact that the species is the first representative of the subgenus to become known from 
<collectingRegion id="E65DDFB0BC4CFFB0FCBFFE33FC8727AD" box="[780,855,401,423]" country="Turkey" name="Burdur" pageId="12" pageNumber="1147">Burdur</collectingRegion>
.
</paragraph>
<caption id="70E641DABC4CFFB0FF01FABBFEF12379" ID-DOI="http://doi.org/10.5281/zenodo.5325913" ID-Zenodo-Dep="5325913" httpUri="https://zenodo.org/record/5325913/files/figure.png" pageId="12" pageNumber="1147" startId="12.[178,218,1305,1327]" targetBox="[178,1076,433,1281]" targetPageId="12">
<paragraph id="24261152BC4CFFB0FF01FABBFEF12379" blockId="12.[178,1085,1305,1395]" pageId="12" pageNumber="1147">
<emphasis id="16EDCD40BC4CFFB0FF01FABBFEC62324" bold="true" box="[178,278,1305,1327]" pageId="12" pageNumber="1147">Figs 26-32</emphasis>
: 
<taxonomicName id="E3996AD1BC4CFFB0FE91FABBFE052324" authority="Assing, 2011" authorityName="Assing" authorityYear="2011" box="[290,469,1305,1326]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1147" phylum="Arthropoda" rank="species" species="burdurica" status="sp. nov.">
<emphasis id="16EDCD40BC4CFFB0FE91FABBFE052324" box="[290,469,1305,1326]" italics="true" pageId="12" pageNumber="1147">Geostiba burdurica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DDE703BBC4CFFB0FE68FAB9FDCE2325" box="[475,542,1307,1327]" pageId="12" pageNumber="1147" rank="species">nov.sp.</taxonomicNameLabel>
(26, 29: holotype): (
<emphasis id="16EDCD40BC4CFFB0FD6FFABBFD232324" bold="true" box="[732,755,1305,1326]" pageId="12" pageNumber="1147">26</emphasis>
) male forebody; (
<emphasis id="16EDCD40BC4CFFB0FC29FABBFC612324" bold="true" box="[922,945,1305,1326]" pageId="12" pageNumber="1147">27</emphasis>
) male abdominal apex in lateral view; (
<emphasis id="16EDCD40BC4CFFB0FE28FA92FE62234F" bold="true" box="[411,434,1328,1349]" pageId="12" pageNumber="1147">28</emphasis>
) posterior portion of male abdominal tergite VII in antero-dorsal view; (
<emphasis id="16EDCD40BC4CFFB0FF0AFAE4FF3F2351" bold="true" box="[185,239,1350,1371]" pageId="12" pageNumber="1147">29-30</emphasis>
) median lobe of aedeagus in lateral view; (
<emphasis id="16EDCD40BC4CFFB0FDCEFAE4FD622351" bold="true" box="[637,690,1350,1371]" pageId="12" pageNumber="1147">31-32</emphasis>
) spermatheca. Scale bars: 26: 0.5 mm; 27- 32: 0.1 mm.
</paragraph>
</caption>
<paragraph id="24261152BC4CFFB0FF01FA22FD4323C4" blockId="12.[178,1085,1408,1486]" pageId="12" pageNumber="1147">
C o m p a r a t i v e n o t e s: Using the key in 
<bibRefCitation id="40086CA3BC4CFFB0FD74FA22FCB2239C" author="ASSING V" box="[711,866,1408,1430]" pageId="12" pageNumber="1147" pagination="1191 - 1246" refId="ref12400" refString="ASSING V. (2009): A revision of Geostiba of the Western Palaearctic region. XIX. New species from Turkey and Iran and additional records, with an updated key and catalogue of the species of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae: Aleocharinae). - Linzer biologische Beitrage 41 (2): 1191 - 1246." type="journal article" year="2009">ASSING (2009)</bibRefCitation>
, 
<taxonomicName id="E3996AD1BC4CFFB0FCDCFA23FC25239F" authorityName="Assing" authorityYear="2011" box="[879,1013,1408,1430]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1147" phylum="Arthropoda" rank="species" species="burdurica">
<emphasis id="16EDCD40BC4CFFB0FCDCFA23FC25239F" box="[879,1013,1408,1430]" italics="true" pageId="12" pageNumber="1147">G. burdurica</emphasis>
</taxonomicName>
would key out at couplets 91-93. For a supplement to this key, which accounts for the new species, see the following section on 
<taxonomicName id="E3996AD1BC4CFFB0FD99FA1BFD5E23C7" authorityName="Assing" authorityYear="2011" box="[554,654,1464,1486]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1147" phylum="Arthropoda" rank="species" species="akiana">
<emphasis id="16EDCD40BC4CFFB0FD99FA1BFD5E23C7" box="[554,654,1464,1486]" italics="true" pageId="12" pageNumber="1147">G. akiana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="24261152BC4CFFB0FF01FA7FFC7D204C" blockId="12.[178,1085,1500,1606]" pageId="12" pageNumber="1147">
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: Thetypelocalityissituatedin the Katrancık Dağı to the southeast of 
<collectingRegion id="E65DDFB0BC4CFFB0FDF5FA5AFD5E2004" box="[582,654,1528,1550]" country="Turkey" name="Burdur" pageId="12" pageNumber="1147">Burdur</collectingRegion>
in 
<collectingRegion id="E65DDFB0BC4CFFB0FD00FA5AFC8F2004" box="[691,863,1528,1550]" country="Turkey" name="Burdur" pageId="12" pageNumber="1147">Burdur province</collectingRegion>
. The specimens were collected in a cypress forest with oak undergrowth at an altitude of approximately 
<quantity id="E361BCB7BC4CFFB0FF01F992FED1204C" box="[178,257,1584,1606]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="12" pageNumber="1147" unit="m" value="1420.0">1420 m</quantity>
, by sifting litter and grass roots near snow (SCHÜLKE, pers. comm).
</paragraph>
</subSubSection>
</treatment>
</document>