Characters | Holotype Range | Mean±SD |
---|---|---|
In % of standard length (SL) | ||
Lateral head length (HL) | 26.5 23.8–26.7 | 25.4±1.06 |
Dorsal head length | 21.6 20.7–22.7 | 21.8±0.73 |
Predorsal length (PDL) | 56.9 53.0–56.9 | 55.3±1.56 |
Preventral length | 60.5 56.3–60.5 | 58.3±1.45 |
Preanal length | 76.2 71.7–76.2 | 73.8±1.45 |
Pre-anus length | 71.5 68.6–71.5 | 70.5±1.30 |
Distance from pectoral fin to pelvic fin origin | 32.5 29.6–34.1 | 31.9±1.92 |
Caudal peduncle length (CPL) | 18.1 18.1–22.6 | 20.0±2.05 |
Caudal peduncle depth (CPD) | 7.8 6.7–9.1 | 7.9±0.97 |
Body depth (at dorsal origin) (BD) | 16.0 12.4–17.1 | 14.5±1.75 |
Body depth (at nape) | 14.1 13.0–14.2 | 13.7±0.41 |
Body width (at dorsal origin) | 11.6 9.4–12.1 | 11.0±1.01 |
Head width (maximum) | 16.2 14.5–16.4 | 15.2±0.85 |
Head height (at eye) | 12.7 8.2–14.4 | 11.6±2.13 |
Height of dorsal fin | 17.2 16.9–19.0 | 18.0±0.83 |
Length of pectoral fin | 18.8 18.8–22.0 | 21.2±1.18 |
Length of pelvic fin | 17.0 14.4–17.6 | 16.2±1.10 |
Length of anal fin | 18.0 17.3–19.4 | 18.3±0.77 |
Length of caudal fin | 23.0 22.5–24.7 | 23.2±0.78 |
Length of median caudal ray | 17.7 17.7–21.5 | 20.3±1.39 |
In % of dorsal head length | ||
Snout length | 38.0 38.0–48.1 | 43.1±3.36 |
Eye diameter (ED) | 18.2 18.2–27.5 | 22.5±3.02 |
Interorbital width | 26.7 23.2–35.1 | 28.0±5.18 |
Head width (maximum) | 74.8 65.8–74.8 | 69.6±3.65 |
Head width (at nare) | 50.4 30.9–50.4 | 45.8±7.41 |
Head height (at eye) | 59.0 36.3–63.8 | 53.1±10.00 |
Outer rostral barbel length | 45.6 28.8–50.9 | 43.4±7.66 |
Maxillary barbel length | 45.8 34.5–53.4 | 41.9±7.27 |
In % of lateral head length (HL) | ||
Snout length | 31.0 31.0–41.4 | 37.1±3.66 |
Eye diameter (ED) | 14.8 14.8–23.7 | 19.4±2.89 |
Interorbital width | 21.8 19.6–30.7 | 24.2±4.97 |
In % of head width (maximum) | ||
Mouth width | 52.3 37.2–52.3 | 45.2±5.46 |
Body width(at dorsal origin) | 71.9 64.5–82.9 | 72.6±5.90 |
Body width (at anal origin) | 54.9 44.2–55.8 | 49.0±5.11 |
Caudal peduncle width (CPW) | 34.3 30.7–37.3 | 34.3±2.39 |
Caudal peduncle: length/depth (CPL/CPD) | 2.3 2.3 – 3.3 | 2.6±0.40 |
Caudal peduncle: width/depth (CPW/CPD) | 0.7 0.6 – 0.7 | 0.66±0.04 |
Characteristic |
|
|
|
|
|
---|---|---|---|---|---|
SL (mm) | 36.8–43.4 | 46–130 | 65–143 | 71–121 | 40–94 |
D A | iii, 7 iii, 5 | iii–iv, 7–8 (7) iii–iv, 5 | iv, 7–8 iii–iv, 5 | iii–iv, 7–8 (7) iii, 5 | iii–iv, 7–10 (8) ii–iii, 5–6 |
P | i, 10–11 | i, 10–12 | i, 10–13 | i, 10–11 | i, 9–13 |
V C | i, 7 15–16(15) | i, 7 16–17 (16) | i, 6–8 15–16 (16) | i, 7 14–18 (15–16) | i, 5–8 13–16 |
Gill rakers | 12 | 10–14 | 12–16 | 12–15 | 11–19 |
Vertebrae HL % SL | 4+33–35 23.8–26.7 | 4+34–36 20.0–25.6 | 4+37–38 20.4–25.0 | 4+36–38 16.7–25.6 | 4+33–38(35) 18.2–27.8 |
BD % SL | 12.4–17.1 | 13.9–22.7 | 15.2–19.6 | 11.5–18.2 | 11.0–22.7 |
CPL % SL CPL/CPD | 18.1–22.6 2.3–3.3 | 14.3–23.3 1.5–2.8 | 20.0–25.0 3.1–5.5 | 23.3–29.4 3.9–9.1 | 21.3–30.3 3.2–7.3 |
CPW <CPD | Yes | Yes | No | No | No |
PDL % SL Position of pelvic fin origin compared with dorsal fin origin Lips papillae developed | 53.0–56.9 Below 2nd–3rd branched dorsal ray Yes | 50–56 Below 3rd–7th branched dor- sal ray No | 50–54 Below 1st – 2nd branched dor- sal ray Yes | 44–50 Below the last unbranched, 1st –2nd branched dorsal ray Yes | 47–52 Opposite, or 1st – 2nd branched dorsal ray Yes |
Intestine, anterior bend reaching which part of the U-shaped stomachic dilatation | 2 coils, mid part | 3–5 coils, ante- rior part | 2 coils, anterior part | 2 coils, anterior part | 2 coils, mid part |
Shape and structure of posterior part of air bladder | Ovoid, with a long tube | Ovoid, with a short tube | Ovoid, with a long tube | Ovoid, with a long tube | Long bag shaped, no tube, |
Caudal fin | Slightly emargin- ate | Slightly emar- ginate | Lunate | Emarginate | Lunate |
continued. |
Characteristic SL (mm) |
|
|
|
|
|
---|---|---|---|---|---|
D | iii, 8 | iv, 8 | iii, 7–8 | iv, 6–7 (7) | iii–iv, 6–7 (7) |
A P | iii, 5 i, 12 | iii, 5 i, 10–11 | ii, 5 i, 10 | iii, 5 i, 9–10 | iv, 5–6 (5) i, 11–12 |
V | i, 7 | i, 7 | i, 6–7 | i, 6–7 | i, 8–9 |
C | 15–16 | 16 | 13–17 (16) | ||
Gill rakers Vertebrae | 12–14 4+35–38 | 4+38 | 15–19(16) 4+36–37 | 8–16 4+37–43 | |
HL % SL | 20 | 17.2–20.4 | 19.2–20.4 | 22.7–32.3 | 18.9–25.6 |
BD % SL | 13.2–14.1 | 10.9–12.5 | 9.3–10.3 | 14.1–27.8 | 12.2–19.6 |
CPL % SL | 22.2–23.3 | 28.6–31.3 | 25.6–30.3 | 15.9–25.6 | 17.5–25.6 |
---|---|---|---|---|---|
CPL/CPD | 3.3 | 7.6–13.1 | 6.7–8.7 | 3.0–3.7 | 1.8–2.8 |
CPW <CPD | Yes | No | No | Yes | Yes |
PDL % SL | >50 | 41–44 | 49.2–50 | Slightly <50 | 55–62 |
Position of pelvic fin origin compared with dorsal fin origin | Below 1st –2nd branched dorsal ray | Below 1st–2nd branched dor- sal ray | Below 3rd branched dorsal ray | Below 1st– 2nd branched dorsal ray | Opposite, or ante- rior |
---|---|---|---|---|---|
Lips papillae developed | No | Yes | No | No | No |
Intestine, anterior bend reaching which part of the U-shaped stomachic dilatation | 2 coils | 2 coils, anterior part | 4 coils, ante- rior part | 2–3 coils | 2 coils, posterior part, |
Shape and structure of posterior part of air bladder | Ovoid, with a long tube | Ovoid, with a long tube | Ovoid, with a tube | Spheric, no tube | Long bag shaped, no tube, |
Caudal fin | Slightly emar- ginate | Emarginate | Not clear | Slightly emarginate | Slightly emargin- ate |
continued. |
Characteristic |
|
|
|
|
---|---|---|---|---|
SL (mm) | 47–85 | 64.5–87 | 89–123 | 80–117 |
D | iii, 6–7 (7) | iii–iv, 7 | Iii, 7 | iii, 7–8 (8) |
A | iii, 5 | iii, 5 | ii, 5 | iii, 5–6 (6) |
P | i, 10–11 | i, 10–11 | i, 11–12 | i, 14–15 |
V | i, 7 | i, 6–7 | i, 7 | i, 8–9 |
C | 14–17 | 16–17 (16) | ||
Gill rakers | 10–14 | 13–16 | 10 | 13–15 |
Vertebrae | 4+36–38 | 4+34–35 | 4+37 | 4+38–43 |
HL % SL | 20.8–23.3 | 21.7–25.0 | 23.3–25.0 | 22.7–28.6 |
BD % SL | 14.7–18.2 | 15.9–20.0 | 15.4–16.9 | 15.2–18.5 |
CPL % SL | 20.8–23.3 | 18.9–22.7 | 15.2–15.4 | 13.3–20.0 |
CPL/CPD | 2.3–2.9 | 1.7–2.3 | 1.9–1.94 | 1.4–2.6 |
CPW <CPD | Yes | Yes | Yes | Yes |
PDL % SL | 50–53 | 53–58 | 56–59 | >50 |
Position of pelvic fin origin compared with dorsal fin ori- gin | Below 1st or 2nd branched ray | Opposite, or 1st branched dorsal ray | Opposite | Slightly posterior |
---|---|---|---|---|
Lips papillae developed | No | No | Yes | No |
Intestine, anterior bend reaching which part of the U- shaped stomachic dilatation | >3 coils, anterior part | 2 coils, posterior part | Not clear | 2 coils, posterior part |
Shape and structure of poste- rior part of air bladder | Ovoid, no tube | Long bag or bottle gourd shaped, no tube | Long bag shaped, no tube | Bottle gourd shaped, no tube |
Caudal fin | Truncate or slightly emarginate | Truncate or slightly emarginate | Truncate | Truncate |