Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood
Author
Frolov, Andrey V.
Author
Montreuil, Olivier
Author
Akhmetova, Lilia A.
text
Zootaxa
2016
4207
1
1
93
journal article
37549
10.11646/zootaxa.4207.1.1
cef35771-ff95-4a41-9af6-196dea899a64
1175-5326
200444
F2875582-31E2-496F-AEEF-1D657DD86C33
Madecorphnus aquilonius
Frolov, 2012
(
Figs. 20
A–F)
Madecorphnus aquilonius
Frolov, 2012b
: 162
.
Type
material examined.
Holotype
(
Figs. 20
A–E), male, “
MADAGASCAR
,
Province Diego-Suarez
,
Sakalava Beach
, dwarf littoral forest elev
10 m
,
13–16 May 2001
/
12°15′46″S
,
49°23′51″E
R.Harin'Hala
collector malaise trap
— across sandy trail MA-01-04B-09 / CASENT 8014036 /
HOLOTYPUS
Madecorphnus aquilonius
Frolov 2011
” (
CASC
)
.
FIGURE 20.
Madecorphnus aquilonius
, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F).
Diagnosis.
Madecorphnus aquilonius
can easily be separated from other
Madecorphnus
species by having characteristic internal sac armature consisting of 2 rather large sclerites: a spur-like sclerite and an asymmetrical, claw-shaped sclerite (
Figs. 20
D–E).
Description. Male.
Body length
5.8 mm
. Color uniformly brown (
Fig. 20
A).
Right mandible about 1/2 longer than left, without tooth behind apex. Labrum trapezoidal, with rounded sides, length about 1/6 width (in dorsal view). Frontoclypeus distinctly asymmetrical, apically obtuse, with 4 relatively long setae on the apical margin. Canthus and frontal suture indistinct. Frontoclypeus slightly depressed apicomedially. Head without traces of frontoclypeal suture, finely punctate with minute punctures separated by greater than 4 times their diameter.
Pronotum approximately 1.75 times wider than long, widest medially. Disc of pronotum convex, without any depressions, tubercles, or ridges. Punctation on pronotum as fine as that on head. Margins with relatively wide border, lateral margins with 4 long setae: 1 seta on basal angle, 1 seta approximately in the middle of lateral margin, and 2 setae on the apical angle.
Scutellum triangular, angulate apically, about 1/11 length of elytra.
Elytra convex, with distinct humeral humps, widest at basal third. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra sparsely punctate with relatively large punctures. Epipleura with long, sparse, brown setae. Base of elytron with border connected to first elytral stria.
Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and 2 smaller setae basally. Mesothoracic and metathoracic legs similar in shape to each other, metathoracic legs 1.4 times longer than mesothoracic legs. Longer tibial spur as long as 2 basal tarsomeres in mesothoracic legs and longer than 2 tarsomeres in metathoracic legs.
Parameres with small teeth laterally, narrowly rounded apically in dorsal view and curved downwards in lateral view (
Fig. 20
B). Internal sac with 2 rather large sclerites: a spur-like sclerite and an asymmetrical, claw-shaped sclerite (
Figs. 20
D–E).
Female.
Unknown.
Distribution and habitat.
This species is known from one locality in the northernmost
Madagascar
in a sandy littoral forest of the
Madagascar
dry deciduous forest ecoregion (
Fig. 20
F).