<document id="F2BE2066B590779AEB71D5CCBF735A5E" ID-DOI="10.5852/ejt.2013.37" ID-GBIF-Dataset="9682b45b-5e22-4965-a293-e203660e0ed3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3806770" ID-ZooBank="ADF6394E-77B5-4309-87FE-4097FDE0A3FD" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1588796067459" checkinUser="carolina" docAuthor="Rowson, Ben &amp; Tattersfield, Peter" docDate="2013" docId="4511E41DD83DFFDDFE63FBB7F6E2FD5C" docLanguage="en" docName="ejt-2013-37.pdf.imf" docOrigin="European Journal of Taxonomy 37 (37)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dadagulella meredithae Rowson &amp; Tattersfield 2013, comb. nov." docType="treatment" docVersion="13" lastPageNumber="27" masterDocId="B9289C65D824FFC7FFA4FF80F253FFE7" masterDocTitle="Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies" masterLastPageNumber="46" masterPageNumber="1" pageNumber="26" updateTime="1743733079341" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9AAA53ADDEA2CA0F32B37AEFB9ABF62D">Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies</mods:title>
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<mods:namePart id="0796B34C17A2029B6409AC09320E2216">Rowson, Ben</mods:namePart>
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<mods:affiliation id="EBBC57575D11C9A0F4A9EA64954391F2">Biodiversity &amp; Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP.</mods:affiliation>
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<mods:namePart id="11A0DCA1A16ABF709B0D38AE63026308">Tattersfield, Peter</mods:namePart>
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<mods:affiliation id="F0B3A93058C69E83C53556066B3F44BD">Biodiversity &amp; Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP.</mods:affiliation>
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<mods:title id="DDFDF35A9A76511BBDDF92A42948F8BE">European Journal of Taxonomy</mods:title>
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<mods:date id="77EB3F3EB50404F5E003370EE2EB98C1">2013</mods:date>
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<treatment id="4511E41DD83DFFDDFE63FBB7F6E2FD5C" ID-DOI="http://doi.org/10.5281/zenodo.3815781" ID-GBIF-Taxon="163971583" ID-Zenodo-Dep="3815781" LSID="urn:lsid:plazi:treatment:4511E41DD83DFFDDFE63FBB7F6E2FD5C" httpUri="http://treatment.plazi.org/id/4511E41DD83DFFDDFE63FBB7F6E2FD5C" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" scope_class="Gastropoda" scope_family="Streptaxidae" scope_order="Stylommatophora" scope_superFamily="Streptaxoidea">
<subSubSection id="85A20680D83DFFDEFE63FBB7F63EFBB5" box="[455,1133,1079,1106]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="CD07550BD83DFFDEFE63FBB7F63EFBB5" blockId="25.[455,1133,1079,1143]" box="[455,1133,1079,1106]" pageId="25" pageNumber="26">
<heading id="964FE267D83DFFDEFE63FBB7F63EFBB5" box="[455,1133,1079,1106]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="0AB82E88D83DFFDEFE63FBB7F1B7FBB5" ID-CoL="33XQJ" authority="(van Bruggen, 2000)" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="2000" box="[455,996,1079,1106]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="meredithae" status="comb. nov.">
<emphasis id="FFCC8919D83DFFDEFE63FBB7F0BAFBB5" bold="true" box="[455,745,1079,1106]" italics="true" pageId="25" pageNumber="26">Dadagulella meredithae</emphasis>
(van 
<bibRefCitation id="A92928FAD83DFFDEFC88FBB8F18FFBB5" author="Bruggen A. C. van" box="[812,988,1080,1106]" pageId="25" pageNumber="26" pagination="225 - 235" refId="ref26131" refString="Bruggen A. C. van. 2000. Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malawi. 5. Description of Gulella meredithae spec. nov. Zoologische Mededelingen 74 (15): 225 - 235." type="journal article" year="2000">Bruggen, 2000</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="E4FF3462D83DFFDEFC48FBB8F63EFBB5" box="[1004,1133,1080,1106]" pageId="25" pageNumber="26" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="85A20680D83DFFDEFD01FBDCF1DDFB90" box="[677,910,1116,1143]" pageId="25" pageNumber="26" type="description">
<paragraph id="CD07550BD83DFFDEFD01FBDCF1DDFB90" blockId="25.[455,1133,1079,1143]" box="[677,910,1116,1143]" pageId="25" pageNumber="26">
<figureCitation id="5583498ED83DFFDEFD01FBDCF0ACFB90" box="[677,767,1116,1143]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="25" pageNumber="26">Figs 37</figureCitation>
, 
<figureCitation id="5583498ED83DFFDEFCAFFBDDF17FFB90" box="[779,812,1117,1143]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="25" pageNumber="26">84</figureCitation>
; 
<tableCitation id="803A60B0D83DFFDEFC93FBDCF1DDFB90" box="[823,910,1116,1143]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetBox="[202,1925,710,1733]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="25" pageNumber="26" tableDoi="http://doi.org/10.5281/zenodo.15137075" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
</paragraph>
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<subSubSection id="85A20680D83DFFDEFF19FB20F152FAC6" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="CD07550BD83DFFDEFF19FB20F134FB5D" blockId="25.[189,871,1183,1210]" box="[189,871,1183,1210]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="EDA87225D83DFFDEFF19FB20F134FB5D" box="[189,871,1183,1210]" pageId="25" pageNumber="26">
<treatmentCitation id="4C19731AD83DFFDEFF19FB20F0A9FB5E" author="Bruggen A. C. van" box="[189,762,1183,1210]" page="232" pageId="25" pageNumber="26" year="2000">
<taxonomicName id="0AB82E88D83DFFDEFF19FB20F0A9FB5E" ID-CoL="6L5L4" authority="van Bruggen, 2000: 226 - 232" authorityName="van Bruggen" authorityPageNumber="226 - 232" authorityYear="2000" box="[189,762,1183,1210]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="meredithae">
<emphasis id="FFCC8919D83DFFDEFF19FB20F3CCFB5D" box="[189,415,1184,1210]" italics="true" pageId="25" pageNumber="26">Gulella meredithae</emphasis>
van 
<bibRefCitation id="A92928FAD83DFFDEFE7CFB20F0A9FB5E" author="Bruggen A. C. van" box="[472,762,1183,1210]" pageId="25" pageNumber="26" pagination="225 - 235" refId="ref26131" refString="Bruggen A. C. van. 2000. Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malawi. 5. Description of Gulella meredithae spec. nov. Zoologische Mededelingen 74 (15): 225 - 235." type="journal article" year="2000">Bruggen, 2000: 226-232</bibRefCitation>
</taxonomicName>
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, figs 1-7.
</treatmentCitationGroup>
</paragraph>
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<paragraph id="CD07550BD83DFFDEFF19FB63F6E5FB1A" blockId="25.[189,1206,1250,1314]" box="[189,1206,1250,1277]" pageId="25" pageNumber="26">
“…an as yet unidentified species from 
<collectingCountry id="B5AF159BD83DFFDEFD21FB62F0B2FB1B" box="[645,737,1250,1276]" name="Malawi" pageId="25" pageNumber="26">Malawi</collectingCountry>
” – van 
<treatmentCitation id="4C19731AD83DFFDEFC9FFB63F6E2FB1A" box="[827,1201,1250,1277]" pageId="25" pageNumber="26">
<bibRefCitation id="A92928FAD83DFFDEFC9FFB63F6E2FB1A" author="Bruggen A. C. van &amp; Meredith H. M." box="[827,1201,1250,1277]" pageId="25" pageNumber="26" pagination="156 - 171" refId="ref26267" refString="Bruggen A. C. van &amp; Meredith H. M. 1984. A preliminary analysis of the land molluscs of Malawi. In: Solem A. &amp; Bruggen A. C. van (eds) World-wide Snails: Biogeographical Studies on non-marine Mollusca: 156 - 171. Backhuys, Leiden, the Netherlands." type="book chapter" year="1984">
Bruggen &amp; Meredith 
<date id="B90673CBD83DFFDEFB9CFB63F624FB1A" box="[1080,1143,1251,1277]" pageId="25" pageNumber="26" value="1984">1984</date>
: 165
</bibRefCitation>
</treatmentCitation>
.
</paragraph>
<paragraph id="CD07550BD83DFFDEFF19FA87F152FAC6" blockId="25.[189,1206,1250,1314]" box="[189,769,1287,1314]" pageId="25" pageNumber="26">
<emphasis id="FFCC8919D83DFFDEFF19FA87F3E5FAC6" box="[189,438,1287,1313]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="0AB82E88D83DFFDEFF19FA87F3CCFAC6" ID-CoL="6L5L4" authorityName="van Bruggen" authorityYear="2000" box="[189,415,1287,1313]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="meredithae">Gulella meredithae</taxonomicName>
–
</emphasis>
<treatmentCitation id="4C19731AD83DFFDEFE18FA88F0AEFAC6" author="Rowson B. &amp; Lange C. N." box="[444,765,1287,1314]" page="31" pageId="25" pageNumber="26" year="2007">
<bibRefCitation id="A92928FAD83DFFDEFE18FA88F0AEFAC6" author="Rowson B. &amp; Lange C. N." box="[444,765,1287,1314]" pageId="25" pageNumber="26" pagination="21 - 32" refId="ref27291" refString="Rowson B. &amp; Lange C. N. 2007. Two new species of Gulella (Mollusca: Pulmonata: Streptaxidae) from the Taita Hills, Kenya. African Invertebrates 48 (2): 21 - 32." type="journal article" year="2007">
Rowson &amp; Lange 
<date id="B90673CBD83DFFDEFD37FA87F081FAC6" box="[659,722,1287,1313]" pageId="25" pageNumber="26" value="2007">2007</date>
: 31
</bibRefCitation>
</treatmentCitation>
.
</paragraph>
</treatmentCitationGroup>
</subSubSection>
<subSubSection id="85A20680D83DFFDEFF19FACBF357F9CA" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="CD07550BD83DFFDEFF19FACBF3BFFA83" blockId="25.[189,492,1354,1381]" box="[189,492,1354,1381]" pageId="25" pageNumber="26">
<heading id="964FE267D83DFFDEFF19FACBF3BFFA83" bold="true" box="[189,492,1354,1381]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="FFCC8919D83DFFDEFF19FACBF3BFFA83" bold="true" box="[189,492,1354,1381]" pageId="25" pageNumber="26">
<typeStatus id="1203EBA9D83DFFDEFF19FACBF2A9FA82" box="[189,250,1355,1381]" pageId="25" pageNumber="26">Type</typeStatus>
material examined
</emphasis>
</heading>
</paragraph>
<paragraph id="CD07550BD83DFFDEFF19FAFBF633FA5E" blockId="25.[189,1399,1402,1466]" pageId="25" pageNumber="26">
<materialsCitation id="7DD05F56D83DFFDEFF19FAFBF633FA5E" ID-GBIF-Occurrence="2610647314" collectingDate="1983-09-16" collectionCode="RMNH" collectorName="H. M. Meredith" country="Malawi" elevation="2100" latitude="-10.6" location="Nyika National Park" longLatPrecision="7789" longitude="33.8" pageId="25" pageNumber="26" specimenCode="RMNH.59399" specimenCount="1" specimenCount-adult="1" stateProvince="Rumphi District" typeStatus="holotype">
<collectingCountry id="B5AF159BD83DFFDEFF19FAFBF368FA72" box="[189,315,1403,1429]" name="Malawi" pageId="25" pageNumber="26">MALAWI</collectingCountry>
: 
<typeStatus id="1203EBA9D83DFFDEFEEFFAFAF3E1FA73" box="[331,434,1402,1428]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
<specimenCode id="9D1EFD70D83DFFDEFE1BFAFAF03FFA72" box="[447,620,1402,1429]" pageId="25" pageNumber="26">RMNH.59399</specimenCode>
: 
<specimenCount id="DBBE9E82D83DFFDEFDD8FAFBF0EFFA73" box="[636,700,1402,1429]" pageId="25" pageNumber="26" type="adult">1 ad.</specimenCount>
, 
<location id="C86703D0D83DFFDEFD6AFAFAF199FA73" LSID="urn:lsid:plazi:treatment:4511E41DD83DFFDDFE63FBB7F6E2FD5C:C86703D0D83DFFDEFD6AFAFAF199FA73" box="[718,970,1402,1428]" country="Malawi" latitude="-10.6" longLatPrecision="7789" longitude="33.8" name="Nyika National Park" pageId="25" pageNumber="26" stateProvince="Rumphi District">Nyika National Park</location>
(approx. 
<geoCoordinate id="A88C33CCD83DFFDEFBE2FAFAF6CBFA73" box="[1094,1176,1402,1428]" degrees="10.6" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="5555" value="-10.6">10.6°S</geoCoordinate>
, 
<geoCoordinate id="A88C33CCD83DFFDEFB0DFAFBF6ADFA72" box="[1193,1278,1403,1429]" degrees="33.8" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="5555" value="33.8">33.8°E</geoCoordinate>
), 
<collectingRegion id="0F7C9BE9D83DFFDEFABCFAFAF34BFA5E" country="Malawi" name="Rumphi" pageId="25" pageNumber="26">Rumphi District</collectingRegion>
, 
<location id="C86703D0D83DFFDEFE80FA1FF39EFA5D" LSID="urn:lsid:plazi:treatment:4511E41DD83DFFDDFE63FBB7F6E2FD5C:C86703D0D83DFFDEFE80FA1FF39EFA5D" box="[292,461,1439,1466]" country="Malawi" latitude="-10.6" longLatPrecision="7789" longitude="33.8" name="Juniper Forest" pageId="25" pageNumber="26" stateProvince="Rumphi District">Juniper Forest</location>
, approx. 
<quantity id="0A40F8EED83DFFDEFD9DFA1FF0C1FA5D" box="[569,658,1439,1466]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="25" pageNumber="26" unit="m" value="2100.0">
<elevation id="4695B238D83DFFDEFD9DFA1FF0C1FA5D" box="[569,658,1439,1466]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="25" pageNumber="26" unit="m" value="2100.0">2100 m</elevation>
</quantity>
alt., leg. 
<collectorName id="604D30DDD83DFFDEFD5BFA20F1E7FA5E" box="[767,948,1439,1466]" pageId="25" pageNumber="26">H.M. Meredith</collectorName>
, 
<date id="B90673CBD83DFFDEFC1BFA1FF60FFA5E" box="[959,1116,1439,1465]" pageId="25" pageNumber="26" value="1983-09-16">
<collectingDate id="A9428A23D83DFFDEFC1BFA1FF60FFA5E" box="[959,1116,1439,1465]" pageId="25" pageNumber="26" value="1983-09-16">16 Sep. 1983</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="CD07550BD83DFFDEFF19FA62F3AAFA1B" blockId="25.[189,505,1506,1532]" box="[189,505,1506,1532]" pageId="25" pageNumber="26">
<heading id="964FE267D83DFFDEFF19FA62F3AAFA1B" bold="true" box="[189,505,1506,1532]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="FFCC8919D83DFFDEFF19FA62F3AAFA1B" bold="true" box="[189,505,1506,1532]" pageId="25" pageNumber="26">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="CD07550BD83DFFDEFF19F993F357F9CA" blockId="25.[189,260,1555,1581]" box="[189,260,1555,1581]" pageId="25" pageNumber="26">None.</paragraph>
</subSubSection>
<subSubSection id="85A20680D83DFFDDFF19F9D5F03CFEAE" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
<paragraph id="CD07550BD83DFFDEFF19F9D5F31DF988" blockId="25.[189,334,1621,1647]" box="[189,334,1621,1647]" pageId="25" pageNumber="26">
<heading id="964FE267D83DFFDEFF19F9D5F31DF988" bold="true" box="[189,334,1621,1647]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="FFCC8919D83DFFDEFF19F9D5F31DF988" bold="true" box="[189,334,1621,1647]" pageId="25" pageNumber="26">Description</emphasis>
</heading>
</paragraph>
<paragraph id="CD07550BD83DFFDDFF19F906F03CFEAE" blockId="25.[189,1399,1669,2027]" lastBlockId="26.[189,1398,267,330]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
SHELL (
<figureCitation id="5583498ED83DFFDEFEB5F905F335F947" box="[273,358,1669,1696]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="25" pageNumber="26">Fig. 37</figureCitation>
). Small (
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2.30 
<emphasis id="FFCC8919D83DFFDEFE5BF906F05AF978" box="[511,521,1670,1695]" italics="true" pageId="25" pageNumber="26">-</emphasis>
3.10 mm
</quantity>
high x 
<quantity id="0A40F8EED83DFFDEFD66F906F135F947" box="[706,870,1669,1696]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="25" pageNumber="26" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">
1.40 
<emphasis id="FFCC8919D83DFFDEFD51F906F0ACF978" box="[757,767,1670,1695]" italics="true" pageId="25" pageNumber="26">-</emphasis>
1.60 mm
</quantity>
wide), of 5.5 
<emphasis id="FFCC8919D83DFFDEFC59F906F654F978" box="[1021,1031,1670,1695]" italics="true" pageId="25" pageNumber="26">-</emphasis>
“&lt;6” whorls. Ovate-acuminate, although spire (spire angle 58 
<emphasis id="FFCC8919D83DFFDEFD87F92BF07EF923" box="[547,557,1707,1732]" italics="true" pageId="25" pageNumber="26">-</emphasis>
77°) less acuminate than in other 
<taxonomicName id="0AB82E88D83DFFDEFC61F92AF60BF923" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[965,1112,1706,1732]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83DFFDEFC61F92AF60BF923" box="[965,1112,1706,1732]" italics="true" pageId="25" pageNumber="26">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4FF3462D83DFFDEFBC5F92BF69AF922" box="[1121,1225,1707,1733]" pageId="25" pageNumber="26" rank="genus">gen. nov</taxonomicNameLabel>
species. Apex rounded (in 
<typeStatus id="1203EBA9D83DFFDEFEF0F94FF3EEF90D" box="[340,445,1743,1770]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
) to weakly pointed. Embryonic whorls smoothly granulate. Later whorls with very fine, very numerous ribs (about 27 per mm on penultimate whorl). Sutures of intermediate depth. Umbilicus closed or nearly so. Peristome incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition 3-fold to 4-fold, consisting of at least: one lamella-like parietal tooth; one slab-like palatal tooth, not forming a parieto-palatal sinus; and a mammillate columellar baffle. Additional teeth limited to a shallow, weak columellar swelling. Juvenile shells with 2-fold to 3-fold dentition: one parietal tooth; one columellar tooth; and usually one basal tooth (termed labral by van 
<bibRefCitation id="A92928FAD83DFFDEFE5BF851F0F9F80C" author="Bruggen A. C. van" box="[511,682,2001,2027]" pageId="25" pageNumber="26" pagination="225 - 235" refId="ref26131" refString="Bruggen A. C. van. 2000. Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malawi. 5. Description of Gulella meredithae spec. nov. Zoologische Mededelingen 74 (15): 225 - 235." type="journal article" year="2000">
Bruggen 
<date id="B90673CBD83DFFDEFDCEF851F0F9F80C" box="[618,682,2001,2027]" pageId="25" pageNumber="26" value="2000">2000</date>
</bibRefCitation>
). Van 
<bibRefCitation id="A92928FAD83DFFDEFD54F851F1FAF80C" author="Bruggen A. C. van" box="[752,937,2001,2027]" pageId="25" pageNumber="26" pagination="225 - 235" refId="ref26131" refString="Bruggen A. C. van. 2000. Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malawi. 5. Description of Gulella meredithae spec. nov. Zoologische Mededelingen 74 (15): 225 - 235." type="journal article" year="2000">
Bruggen (
<date id="B90673CBD83DFFDEFCC7F851F1F2F80C" box="[867,929,2001,2027]" pageId="25" pageNumber="26" value="2000">2000</date>
)
</bibRefCitation>
showed that earlier sets of dentition are visible through the shells of some transparent juveniles, even in the preceding whorls, suggesting slow or no resorbtion. Anatomy unknown.
</paragraph>
</subSubSection>
<subSubSection id="85A20680D83EFFDDFF19FEF2F3DAFDE5" pageId="26" pageNumber="27" type="biology_ecology">
<paragraph id="CD07550BD83EFFDDFF19FEF2F3F6FE6C" blockId="26.[189,421,369,396]" box="[189,421,369,396]" pageId="26" pageNumber="27">
<heading id="964FE267D83EFFDDFF19FEF2F3F6FE6C" bold="true" box="[189,421,369,396]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="FFCC8919D83EFFDDFF19FEF2F3F6FE6C" bold="true" box="[189,421,369,396]" pageId="26" pageNumber="27">Range and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="CD07550BD83EFFDDFF19FE21F3DAFDE5" blockId="26.[189,1399,416,514]" pageId="26" pageNumber="27">
Montane forest (above 
<quantity id="0A40F8EED83EFFDDFE71FE20F063FE5C" box="[469,560,416,443]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="m" value="1500.0">
<date id="B90673CBD83EFFDDFE71FE20F043FE5C" box="[469,528,416,443]" pageId="26" pageNumber="27" value="1500">1500</date>
m
</quantity>
to around 
<quantity id="0A40F8EED83EFFDDFD11FE20F141FE5C" box="[693,786,416,443]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="26" pageNumber="27" unit="m" value="2450.0">
<date id="B90673CBD83EFFDDFD11FE20F0A3FE5C" box="[693,752,416,443]" pageId="26" pageNumber="27" value="2450">2450</date>
m
</quantity>
) in northern and central 
<collectingCountry id="B5AF159BD83EFFDDFB9BFE20F6CFFE5D" box="[1087,1180,416,442]" name="Malawi" pageId="26" pageNumber="27">Malawi</collectingCountry>
, and adjacent part of 
<collectingCountry id="B5AF159BD83EFFDDFF7AFE44F36BFE39" box="[222,312,452,478]" name="Zambia" pageId="26" pageNumber="27">Zambia</collectingCountry>
(Chowo Forest). Van 
<bibRefCitation id="A92928FAD83EFFDDFD9BFE45F0A9FE39" author="Bruggen A. C. van" box="[575,762,452,479]" pageId="26" pageNumber="27" pagination="225 - 235" refId="ref26131" refString="Bruggen A. C. van. 2000. Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malawi. 5. Description of Gulella meredithae spec. nov. Zoologische Mededelingen 74 (15): 225 - 235." type="journal article" year="2000">Bruggen (2000)</bibRefCitation>
suspected it to range into parts of 
<collectingCountry id="B5AF159BD83EFFDDFB32FE44F6ACFE39" box="[1174,1279,452,478]" name="Tanzania" pageId="26" pageNumber="27">Tanzania</collectingCountry>
adjoining northern 
<collectingCountry id="B5AF159BD83EFFDDFE8CFE68F3D6FDE5" box="[296,389,488,514]" name="Malawi" pageId="26" pageNumber="27">Malawi</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="85A20680D83EFFDDFF19FDABF6E2FD5C" pageId="26" pageNumber="27" type="discussion">
<paragraph id="CD07550BD83EFFDDFF19FDABF37CFDA2" blockId="26.[189,303,555,581]" box="[189,303,555,581]" pageId="26" pageNumber="27">
<heading id="964FE267D83EFFDDFF19FDABF37CFDA2" bold="true" box="[189,303,555,581]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="FFCC8919D83EFFDDFF19FDABF37CFDA2" bold="true" box="[189,303,555,581]" pageId="26" pageNumber="27">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="CD07550BD83EFFDDFF19FDD9F6E2FD5C" blockId="26.[189,1399,601,700]" pageId="26" pageNumber="27">
This species is distinctive in its small size, very fine, numerous ribs, and dentition. It differs from 
<taxonomicName id="0AB82E88D83EFFDDFAFEFDDAF35BFD70" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis id="FFCC8919D83EFFDDFAFEFDDAF35BFD70" italics="true" pageId="26" pageNumber="27">D. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4FF3462D83EFFDDFEB4FDFDF3C2FD7F" box="[272,401,637,664]" pageId="26" pageNumber="27" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="FFCC8919D83EFFDDFE3DFDFDF3E8FD70" box="[409,443,637,663]" italics="true" pageId="26" pageNumber="27">s.l.</emphasis>
and 
<taxonomicName id="0AB82E88D83EFFDDFE50FDFEF039FD70" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[500,618,637,663]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
<emphasis id="FFCC8919D83EFFDDFE50FDFEF039FD70" box="[500,618,637,663]" italics="true" pageId="26" pageNumber="27">D. browni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4FF3462D83EFFDDFDD6FDFDF0A7FD7F" box="[626,756,637,664]" pageId="26" pageNumber="27" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="FFCC8919D83EFFDDFD5FFDFDF14DFD70" box="[763,798,637,663]" italics="true" pageId="26" pageNumber="27">s.l.</emphasis>
in the lack of a basal tooth or denticle. The apex is rounded in the 
<typeStatus id="1203EBA9D83EFFDDFECAFD21F386FD5C" box="[366,469,673,699]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
but more conical in 
<typeStatus id="1203EBA9D83EFFDDFD62FD22F165FD5B" box="[710,822,674,700]" pageId="26" pageNumber="27" type="paratype">paratypes</typeStatus>
figured by van 
<bibRefCitation id="A92928FAD83EFFDDFC4BFD22F6FDFD5C" author="Bruggen A. C. van" box="[1007,1198,673,700]" pageId="26" pageNumber="27" pagination="225 - 235" refId="ref26131" refString="Bruggen A. C. van. 2000. Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malawi. 5. Description of Gulella meredithae spec. nov. Zoologische Mededelingen 74 (15): 225 - 235." type="journal article" year="2000">Bruggen (2000)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>