A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea Author Harzhauser, Mathias Author Landau, Bernard text Zootaxa 2016 4210 1 1 178 journal article 37280 10.11646/zootaxa.4210.1.1 e782e07d-76b7-4e9b-ba34-ed3286254ec6 1175-5326 252966 D39416B8-CF85-440B-84C2-D4380BECC4E3 Monteiroconus pervindobonensis (Sacco, 1893) Figs 26 E1–E2 Conus betulinoides Lam. —Hörnes 1851: 16, pl. 1, figs 1a–b [non Kalloconus betulinoides ( Lamarck, 1810 ) ]. Conus ( Dendroconus ) betulinoides Lam. Hoernes & Auinger 1879 : 17 [non Kalloconus betulinoides ( Lamarck, 1810 ) ]. [ C o nus ( Dendroconus ) betulinoides (Lk.) ] varietà pervindobonensis Sacc. Sacco 1893a : 4 [nov. nom. pro Conus betulinoides sensu Hörnes 1851, pl. 1, fig. 1]. Type material. The holotype illustrated in Hörnes (1851, pl. 1, fig. 1) is lost. Studied material. 1 spec. NHMW 1869/0001/0020, 1 spec. NHMW 2010/0004/1574, 2 spec. NHMW 2010 / 0004/1575 Bad Vöslau ( Austria ) ; 1 spec NHMW A989 Immendorf ( Austria ) ; 2 spec. NHMW 1852 /0012/0010, 2 NHMW 1851 /0026/0004 Grund ( Austria ) . Illustrated material. Figs 26 E1–E2: Bad Vöslau ( Austria ): SL: 124.5, mm, MD: 74.0 mm, 1 spec. NHMW 1869 /0001/0020. Revised description. Very large shells with low spire; spire whorls striate, weakly convex with shallow central concavity in some specimens. Subsutural flexure shallow and asymmetrically curved. Shoulder rounded or faintly subangulate, position of maximum diameter close below shoulder. Last whorl elongate, regular conical in dorsal view, ventricose in apertural view; not constricted. Aperture wide, slightly excavated at junction from fasciole and inner lip. Fasciole broad, swollen, demarcated from glossy, inner lip by distinct groove. Growth lines on last whorl below shoulder prosocline and straight. Siphonal canal wide, slightly deflected. No colour pattern preserved. Shell measurements and ratios. n = 5: largest specimen: SL: 124.5 mm , MD: 74.00 mm, mean SL: 101.3 mm (σ = 13.3), mean MD: 62.8 mm (σ = 6.5), spire angle: µ = 135.6° (σ = 7.3°), last whorl angle: µ = 39.2° (σ = 2.6°), LW: µ = 1.61 (σ = 0.08), RD: µ = 0.69 (σ = 0.02), PMD: µ = 0.9 (σ = 0.02), RSH: µ = 0.1 (σ = 0.02), Hörnes (1851, pl. 1, fig. 1) reports a height of 128 mm and a width of 73 mm for the illustrated but lost specimen. Discussion. The specimen of Hörnes (1851, pl. 1, fig. 1) from Steinebrunn was already lost in 1879 ( Hoernes & Auinger, 1879 ) and only a few additional specimens in the NHMW collection are available. Unfortunately, Hörnes (1851) presented only the dorsal view of his specimen. Therefore, the conspecificity with the Pliocene Kalloconus betulinoides ( Lamarck, 1810 ) remained unchallenged. Only Sacco (1893a) proposed the variety name pervindobonensis for this specimen, without discussing specific differences from the type. A re-investigation of the few additional Paratethyan specimens seems to support Sacco’s position. The concave and striate spire whorls, the excavated columella and the deflected siphonal canal of the Vienna Basin specimens differ from the Italian Kalloconus betulinoides as illustrated by Davoli (1972) , Pinna & Spezia (1978) and Chirli (1997) . The shell is broader and club-shaped, the last whorl less elongate and the last spire whorl is narrower. Therefore, we propose to separate the middle Miocene Paratethyan shells as Monteiroconus pervindobonensis (Sacco, 1893) . The generic placement is based on the concave and striate spire whorls. Paleoenvironment. The specimens from Bad Vöslau, Grund and Immendorf were found in clays indicating an offshore environment but transport from shallow habitats cannot be excluded ( Zuschin et al. 2005 , 2006 ). Distribution in Paratethys.? Karpatian (early Miocene): Alpine-Carpathian Foredeep : Niederkreuzstetten ( Austria ) (Harzhauser 2002). Badenian (middle Miocene): Vienna Basin: Gainfarn, Steinebrunn, Bad Vöslau, ( Austria ); Alpine-Carpathian Foredeep : Grund, Immendorf ( Austria ) ( Hoernes & Auinger 1879 );? Korytnica ( Poland ) ( Bałuk 1997 );? Pannonian Basin : Letkés, Márkháza, Mátraverebély, Sámsonháza, Budapest : Rákos ( Hungary ) ( Kovács & Vicián 2013 );? Krka Basin : Gorenje Vrhpolje ( Slovenia ) ( Mikuž 2009 ).