A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea Author Harzhauser, Mathias Author Landau, Bernard text Zootaxa 2016 4210 1 1 178 journal article 37280 10.11646/zootaxa.4210.1.1 e782e07d-76b7-4e9b-ba34-ed3286254ec6 1175-5326 252966 D39416B8-CF85-440B-84C2-D4380BECC4E3 Kalloconus ponderoaustriacus (Sacco, 1893) Figs 3 P, 11G1–G3, 11H1–H3 Conus ( Rhizoconus ) ponderosus Brocc. Var. III—Hoernes & Auinger 1879 : 39 (partim), pl. 5, fig. 6 [non Lautoconus ponderosus ( Brocchi, 1814 ) ]. [ Conus ( Chelyconus) conoponderosus Sacc. ] var. ponderoaustriaca Sacc. Sacco 1893b : 75 [nov. nom. pro Conus ponderosus in Hoernes & Auinger 1879 , pl. 5, fig. 6]. ? Conus ponderosus Brocc. var. Steinabrunnensis Sacco—Friedberg 1911: 58, text-fig. 12 (non pl. 3, fig. 3) [non Lautoconus steinabrunnensis (Sacco, 1893) ]. ? Conus ( Rhizoconus) ponderosus steinabrunnensis (Sacco) Csepreghy-Meznerics 1956 : plate captions, pl. 11, figs 11–12. Conus ( Chelyconus) ponderosus var. ponderoaustriaca (Sacco) —Kojumdgieva in Kojumdgieva & Strachimirov 1960 : 214 , pl. 51, fig. 1. ? Conus ( Chelyconus) ponderosus var. steinabrunnensis (Sacco) —Kojumdgieva in Kojumdgieva & Strachimirov 1960 : 213 , pl. 50, fig. 8 [non Lautoconus steinabrunnensis (Sacco, 1893) ]. Conus ponderosus Brocc. Eremija 1959 : 187 , pl. 1, figs 5-5a [non Lautoconus ponderosus ( Brocchi, 1814 ) ]. Conus ( Rhisoconus) [sic] ponderosus ponderosus Brocchi—Atanacković 1963: 79 , pl. 15, fig. 1 [non Lautoconus ponderosus ( Brocchi, 1814 ) ]. ? Conus ( Dendroconus ) steindachneri Hoernes et Auinger—Atanacković 1963: 79 , pl. 14, figs 6–6b. Conus ( Chelyconus ) ponderosus ponderoaustriacus (Sacco, 1893) Hinculov 1968 : 151 , pl. 38, figs 4a–b. ? Conus ( Chelyconus ) aff. ponderoaustriacus Sacco, 1893 Strausz 1966 : 462 , pl. 69, figs 7–8. Conus ( Chelyconus ) ponderosus ponderoaustriacus (Sacco, 1893) Nicorici & Sagatovici 1973 : 176 , pl. 27, figs 6–7. ? Conus ( Chelyconus ) ponderosus ponederoaustriacus [sic] (Sacco, 1893)— Atanacković 1985 : 177 , pl. 39, figs 10–11. Conus panderosus [sic] ponderoaustriacus (Sacco) Ionesi & Nicorici 1994 : 62 , pl. 5, figs 5, 7. ? Conus ( Chelyconus ) ponderosus Brocchi—Chira & Voia 2001 : 156 , pl. 3, figs 1a–b. Type material. Holotype NHMW 1854 /0035/0041, Lăpugiu de Sus ( Romania ); middle Miocene , Badenian ( Langhian ). Studied material. Holotype and 1 spec. NHMW 1854/0035/0041, 1 spec. NHMW 1854 /0035/0037, Lăpugiu de Sus ( Romania ) ; 2 spec. NHMW 1856 /0050/0109, Gainfarn ( Austria ) ; 5 spec. NHMW 1855 /0045/0358, Gainfarn ( Austria ) ; 9 spec. NHMW 1861 /0033/0005, Bujtur ( Romania ) . Illustrated material. Figs 11 G1–G3, 3P: holotype , Lăpugiu de Sus ( Romania ), SL: 50.4 mm , MD: 27.1 mm , NHMW 1854/0035/0041; Figs 11 H1–H3: Gainfarn ( Austria ), SL: 49.6 mm , MD: 27.5 mm , NHMW 1855/0045/ 0 358.
FIGURE 12. Principal Components Analysis (PCA) of shell ratios of Kalloconus moravicus and Phasmoconus
fuscocingulatus revealing a clear separation. LWA = angle of the last whorl, LW = length width ratio, RD = relative diameter,
RSH = relative height of spire.
Revised description. Medium-sized shells with low conical to broad coeloconoid spire; early spire whorls nearly flat, later whorls weakly convex with last spire whorl often slightly more convex; spire whorls smooth; suture channelled. Subsutural flexure shallow, weakly curved, moderately asymmetrical. Spire passing via a rounded shoulder into conical to weakly ventricose last whorl; position of maximum diameter close below shoulder. Last whorl faintly constricted; prominent to weak spiral grooves may appear on lower third; siphonal canal short; siphonal fasciole weak and indistinct. Aperture moderately narrow, posteriorly narrowing and only weakly broadening anteriorly. Colour pattern poorly preserved, but seems to consist of densely spaced continuous spirals.
Shell measurements and ratios. n = 10 adult specimens: largest specimen: SL: 59.3 mm , MD: 34.1 mm , mean SL: 49.8 mm (σ = 4.6), mean MD: 27.6 mm (σ = 2.9), spire angle: µ = 115.2° (σ = 8.1°), last whorl angle: µ = 34.6° (σ = 2.5°), LW: µ = 1.8 (σ = 0.1), RD: µ = 0.64 (σ = 0.03), PMD: µ = 0.9 (σ = 0.02), RSH: µ = 0.1 (σ = 0.03). Discussion. Differs from Lautoconus steinabrunnensis in the rounded shoulder, convex spire whorls, lower spire and the higher position of the maximum diameter. Its aperture is narrower especially in the posterior part. Paleoenvironment. Shallow marine environments; at least at Gainfarn, the species was associated with seagrass. Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin : Gainfarn ( Austria ) ( Hoernes & Auinger 1879 ); Transylvanian Basin : Lăpugiu de Sus ( Romania ) ( Hoernes & Auinger 1879 ); Pannonian Basin: Hidas, Szob ( Hungary ) (Csepreghy-Meznerics 1950, 1956; Strausz 1966 ); Zârand Basin : Minişul de Sus ( Romania ) ( Nicorici & Sagatovici 1973 ); Caransebeş-Mehadia Basin: Valea Satului ( Romania ) ( Hinculov 1968 ); Buzău Basin : Valea Muscelului ( Romania ) ( Ionesi & Nicorici 1994 ); Dacian Basin : Dobruša ( Bulgaria ) ( Kojumdgieva & Strachimirov 1960 ); Banja Luka Basin : Jazovac ( Bosnia and Herzegovina ) ( Atanacković 1985 ); Karlovac-Glina Basin: Glina ( Croatia ) ( Eremija 1959 ).