<document id="72EE2FF1A3BA656B6E9C170AB48F1F18" ID-DOI="10.5281/zenodo.5378363" ID-ISSN="1638-9395" ID-Zenodo-Dep="5378363" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630284497952" checkinUser="marcus" docAuthor="Rage, Jean-Claude &amp; Bailon, Salvador" docDate="2005" docId="03F57B1BFFCEFFE2FEFE52C3FF77F73E" docLanguage="en" docName="Geodiversitas.2005.27.3.413-441.pdf" docOrigin="Geodiversitas 27 (3)" docStyle="DocumentStyle:94524207C6714CF92F879FE43E306202.3:Geodiversitas.2000-2008.journal_article.type2" docStyleId="94524207C6714CF92F879FE43E306202" docStyleName="Geodiversitas.2000-2008.journal_article.type2" docStyleVersion="3" docTitle="Ophisaurus undetermined" docType="treatment" docVersion="3" lastPageNumber="425" masterDocId="FFCC0363FFC2FFEEFFD4532FFFCBF27B" masterDocTitle="Amphibians and squamate reptiles from the late early Miocene (MN 4) of Béon 1 (Montréal-du-Gers, southwestern France)" masterLastPageNumber="441" masterPageNumber="413" pageNumber="425" updateTime="1699214189119" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="6D7C6C6DF74A5F716B5FCBE3B0B80C45">Amphibians and squamate reptiles from the late early Miocene (MN 4) of Béon 1 (Montréal-du-Gers, southwestern France)</mods:title>
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<mods:namePart id="1D08023B55F0E9E129E519946A1E0757">Rage, Jean-Claude</mods:namePart>
<mods:affiliation id="6E0FD1A91CC06C5FB0CFEC45F55B38AA">UMR CNRS 5143, Département Histoire de la Terre, Muséum national d’Histoire naturelle, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) jcrage @ mnhn. fr</mods:affiliation>
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<mods:namePart id="CAF274577C0112391AE64877C9778D92">Bailon, Salvador</mods:namePart>
<mods:affiliation id="6B820AC86A1B0538C3C8F210650D7631">UMR CNRS 6569, Laboratoire départemental de Préhistoire du Lazaret, 33 bis boulevard Franck Pilatte, F- 06300 Nice (France) sbailon @ lazaret. unice. fr</mods:affiliation>
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<mods:title id="F7D58A5EC9B4B9882D139F2FCBC9DCFC">Geodiversitas</mods:title>
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<mods:part id="0E18E88B146ED738C4D452DA7EC89D68">
<mods:date id="E9A0B4E80BD7BC884D1925F84D5AA714">2005</mods:date>
<mods:detail id="6216E0A30FE0127386AB86982F62E1D0" type="volume">
<mods:number id="D34B02E0A78E3439BDF04EFF891FD2E8">27</mods:number>
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<mods:number id="C12F3C3A6435790544187A8FAD386159">3</mods:number>
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<mods:start id="4E0686A80B6B4166E7D47C65ECE52FF1">413</mods:start>
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<mods:identifier id="B065BB944532DA91227D9365A32047AB" type="DOI">10.5281/zenodo.5378363</mods:identifier>
<mods:identifier id="FAA033EA5EE18B25DF3CEB7A10D145C1" type="ISSN">1638-9395</mods:identifier>
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<subSubSection id="C3469986FFCEFFE2FEFE52C3FE66F05E" pageId="12" pageNumber="425" type="nomenclature">
<paragraph id="8BE3CA0DFFCEFFE2FEFE52C3FE66F05E" blockId="12.[298,461,492,549]" pageId="12" pageNumber="425">
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<taxonomicName id="4C5CB18EFFCEFFE2FEFE52C3FE06F07E" box="[298,461,492,517]" class="Reptilia" family="Anguidae" genus="Ophisaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="425" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B928161FFFCEFFE2FEFE52C3FE6CF07E" bold="true" box="[298,423,492,517]" italics="true" pageId="12" pageNumber="425">Ophisaurus</emphasis>
sp.
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(
<figureCitation id="1367D688FFCEFFE2FE855123FE6EF05E" box="[337,421,524,549]" captionStart="FIG" captionStartId="11.[113,124,1332,1349]" captionTargetBox="[119,1202,209,1290]" captionTargetId="figure-326@11.[98,1225,195,1304]" captionTargetPageId="11" captionText="FIG. 4. — A, B, Pseudopus laurillardi (Lartet, 1851); A, right dentary (Béon 2004 LT 39); B, posterior part of a braincase (Béon 2004 LT 38); C, Ophisaurus sp., left dentary (Béon 2004 LT 49). Dorsal (d), left lateral (l), medial (m), and right lateral (r) views. Abbreviations: dr, dorsal ridge; pms, posteromedial surface of paroccipital process; vb, ventral border of trigeminal notch. Scale bars: A, B, 5 mm; C, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5378371" httpUri="https://zenodo.org/record/5378371/files/figure.png" pageId="12" pageNumber="425">Fig. 4C</figureCitation>
)
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</paragraph>
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<subSubSection id="C3469986FFCEFFE2FFA55169FF52F0D2" pageId="12" pageNumber="425" type="materials_examined">
<paragraph id="8BE3CA0DFFCEFFE2FFA55169FF52F0D2" blockId="12.[113,646,582,681]" pageId="12" pageNumber="425">MATERIAL EXAMINED. — 1 fragmentary left dentary (Béon 2004 LT 49), and perhaps 1 pterygoid (Béon 2004 LT 50) and 20 vertebrae (Béon 2004 LT 51, 52).</paragraph>
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<subSubSection id="C3469986FFCEFFE2FFA551E3FF77F73E" pageId="12" pageNumber="425" type="discussion">
<paragraph id="8BE3CA0DFFCEFFE2FFA551E3FF3AF09F" blockId="12.[113,646,716,1349]" box="[113,241,716,741]" pageId="12" pageNumber="425">COMMENTS</paragraph>
<paragraph id="8BE3CA0DFFCEFFE2FFA551C3FEC8F63E" blockId="12.[113,646,716,1349]" pageId="12" pageNumber="425">
On the dentary, the subdental shelf and alveolar surface of the parapet form a single, inclined surface; as a result, the implantation of teeth is subpleurodont. The 
<emphasis id="B928161FFFCEFFE2FEE85063FDD2F11E" box="[316,537,844,869]" italics="true" pageId="12" pageNumber="425">lamina horizontalis</emphasis>
does not project medial to the bases of teeth. The teeth are conical; their apices are pointed, slightly recurved and not striated. Such teeth are reminiscent of those of 
<taxonomicName id="4C5CB18EFFCEFFE2FF1A50E3FE8FF19E" authorityName="Daudin" authorityYear="1803" box="[206,324,972,997]" class="Reptilia" family="Anguidae" genus="Ophisaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="425" phylum="Chordata" rank="genus">
<emphasis id="B928161FFFCEFFE2FF1A50E3FE8FF19E" box="[206,324,972,997]" italics="true" pageId="12" pageNumber="425">Ophisaurus</emphasis>
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(
<emphasis id="B928161FFFCEFFE2FE8150E3FE42F19E" box="[341,393,972,997]" italics="true" pageId="12" pageNumber="425">sensu</emphasis>
<bibRefCitation id="EFCDB7FCFFCEFFE2FE4550E3FDF7F19E" author="KLEMBARA J." box="[401,572,972,997]" pageId="12" pageNumber="425" pagination="163 - 169" refId="ref18220" refString="KLEMBARA J. 1979. - Neue Funde der Gattungen Ophisaurus und Anguis (Squamata, Reptilia) aus dem Untermiozan Westbohmens (CSSR). VestnIk Ustredniho ustavu geologickeho 54: 163 - 169." type="journal article" year="1979">Klembara 1979</bibRefCitation>
, 
<bibRefCitation id="EFCDB7FCFFCEFFE2FD9350E3FD4AF19E" author="KLEMBARA J." box="[583,641,972,997]" pageId="12" pageNumber="425" pagination="121 - 168" refId="ref18256" refString="KLEMBARA J. 1981. - Beitrag zur Kenntniss der Subfamilie Anguinae (Reptilia, Anguidae). Acta Universitatis Carolinae-Geologica 2: 121 - 168." type="journal article" year="1981">1981</bibRefCitation>
; i.e. 
<taxonomicName id="4C5CB18EFFCEFFE2FF7250C3FE6DF67E" authority="Gray, 1853" authorityName="Gray" authorityYear="1853" box="[166,422,1004,1029]" class="Reptilia" family="Anguidae" genus="Dopasia" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="425" phylum="Chordata" rank="genus">
<emphasis id="B928161FFFCEFFE2FF7250C3FECDF67E" box="[166,262,1004,1029]" italics="true" pageId="12" pageNumber="425">Dopasia</emphasis>
Gray, 1853
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included) of the European 
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. But a more precise identification is impossible.
</paragraph>
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One pterygoid and 20 vertebrae or so, that display the anguine morphology, are tentatively assigned to this taxon on the basis of size consistency. Moreover, the centrum is less widened anteriorly and the ventral surface more convex than in 
<taxonomicName id="4C5CB18EFFCEFFE2FEA657C3FE15F77E" authorityName="Merrem" authorityYear="1820" box="[370,478,1260,1285]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="425" phylum="Chordata" rank="genus">
<emphasis id="B928161FFFCEFFE2FEA657C3FE15F77E" box="[370,478,1260,1285]" italics="true" pageId="12" pageNumber="425">Pseudopus</emphasis>
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, and the subcentral ridges are slightly concave in ventral aspect.
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