<document id="76169FBECBF093A51C318B5C574BA68F" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia &amp; Ahrens, Dirk" docDate="2017" docId="03CB6967FF8A385A3887FF329339F9E7" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla Erichson 1847" docType="treatment" docVersion="2" lastPageNumber="335" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="335" updateTime="1738712560284" updateUser="ExternalLinkService">
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<mods:title id="27907EF55A4F6B6E8FDB9A21AAFEAE4A">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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<mods:namePart id="A2AFAC351EA499ECA391BAC84C258786">Eberle, Jonas</mods:namePart>
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<mods:namePart id="68C0DBEEA3ED01A5FA4A7ED556397D8D">Beckett, Marc</mods:namePart>
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<mods:namePart id="6A918F2E12E5B259FAF1C3DA570BF996">Ozgul-Siemund, Alev</mods:namePart>
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<mods:namePart id="062AA9DD179166B247FF08ED6A02406B">Frings, Jujina</mods:namePart>
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<heading id="D0956F1DFF8A385A3887FF3294F7FF2D" box="[955,1425,128,155]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="335" reason="2">
NINETEEN NEW 
<taxonomicName id="4C62A3F2FF8A385A3FAFFF329427FF2C" ID-CoL="6SH6" authorityName="Erichson" authorityYear="1847" box="[1171,1345,133,154]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B9160463FF8A385A3FAFFF329427FF2C" box="[1171,1345,133,154]" italics="true" pageId="14" pageNumber="335">PLEOPHYLLA</emphasis>
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335
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</paragraph>
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<caption id="DF1D88F9FF8A385A3BACFC2A9299FC7B" ID-DOI="http://doi.org/10.5281/zenodo.14807097" ID-Zenodo-Dep="14807097" httpUri="https://zenodo.org/record/14807097/files/figure.png" pageId="14" pageNumber="335" startId="14.[144,223,925,944]" targetBox="[144,1428,192,893]" targetPageId="14" targetType="figure">
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<emphasis id="B9160463FF8A385A3BACFC2A919CFC06" bold="true" box="[144,250,925,944]" pageId="14" pageNumber="335">Figure 7.</emphasis>
Distribution of 
<taxonomicName id="4C62A3F2FF8A385A3A96FC2A930FFC06" box="[426,617,925,944]" pageId="14" pageNumber="335">
<emphasis id="B9160463FF8A385A3A96FC2A937EFC06" box="[426,536,925,944]" italics="true" pageId="14" pageNumber="335">Pleophylla</emphasis>
species
</taxonomicName>
(left side) and absence records (right side), represented by sampling localities of all other sampled 
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species without records of 
<taxonomicName id="4C62A3F2FF8A385A383AFC0D9212FC7B" authorityName="Erichson" authorityYear="1847" box="[774,884,954,973]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B9160463FF8A385A383AFC0D9212FC7B" box="[774,884,954,973]" italics="true" pageId="14" pageNumber="335">Pleophylla</emphasis>
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(white dots).
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metatarsomere as long as subsequent one. Aedeagus: 
<figureCitation id="1359C4F4FF8A385A3BACFBA99182FB85" box="[144,228,1054,1076]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="14" pageNumber="335">Fig. 2Y</figureCitation>
<emphasis id="B9160463FF8A385A3BD8FB969196FB85" box="[228,240,1057,1075]" italics="true" pageId="14" pageNumber="335">–</emphasis>
Aa. Habitus: 
<figureCitation id="1359C4F4FF8A385A3AB5FBA990ADFB82" box="[393,459,1054,1076]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="14" pageNumber="335">Fig. 2</figureCitation>
Ab.
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<emphasis id="B9160463FF8A385A3BACFBFA936AFBD4" box="[144,524,1100,1122]" italics="true" pageId="14" pageNumber="335">
Diagnosis: 
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is similar to 
<taxonomicName id="4C62A3F2FF8A385A3BACFBDC9023FB36" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[144,325,1131,1152]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="species" species="burundiensis" status="sp. nov.">
<emphasis id="B9160463FF8A385A3BACFBDC9023FB36" box="[144,325,1131,1152]" italics="true" pageId="14" pageNumber="335">P. burundiensis</emphasis>
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, but it can be distinguished by the basal lobe of the left paramere, which is produced beyond the distal half of the paramere.
</paragraph>
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<emphasis id="B9160463FF8A385A3BACFB6D9065FB59" box="[144,259,1242,1263]" italics="true" pageId="14" pageNumber="335">Variation:</emphasis>
Body length: 
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8.8 
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10.6 mm
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, elytral length: 
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6.9 
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8.1 mm
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, maximum width: 
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4.7 
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5.6 mm
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. Density of dark spots may vary slightly, but never completely without spots. Female: antennal club of female straight, composed of five antennomeres, slightly longer than the remaining antennomeres combined; sixth antennomere short, slightly shorter than half of club length. Eyes smaller than in male: ratio of eye diameter/interocular width: 0.52.
</paragraph>
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<emphasis id="B9160463FF8A385A3BACFA49906BF9A5" box="[144,269,1534,1555]" italics="true" pageId="14" pageNumber="335">Etymology:</emphasis>
The new species is named 
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‘ 
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’ following its occurrence in the eastern 
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(Virunga Mountains).
</paragraph>
</subSubSection>
</treatment>
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