Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae) Author Ballantyne, Lesley A. Author Lambkin, Christine text Zootaxa 2009 2009-02-04 1997 1 188 journal article 1175­5334 Pygatyphella limbatifusca sp. n. ( Figs 329, 338, 341 , 362, 363, 370, 371 ) Holotype . Male. SOLOMON ISLANDS : Makira Pr., San Cristobal : 10.35S , 161.30E , Manipwena , Magoha River , 13.viii.1960 , COB ( BPBM ). Paratypes (26). Collector is COB. SOLOMON ISLANDS : Makira Pr. , San Cristobal : 10.35S , 161.30E , Manipwena , Magoha River , 13.viii.1960 , 6 males , 3 females ; Kira Kira light trap, 15.viii.1960 , male; Napagiwae , 20.viii.1960 , 5 males ; Wugiroga , 10.viii.1960 , male; Bweinaniawarikiapu , 11–12.viii.1960 , 7 males , 4 females ( BPBM ) . Diagnosis . Part of the Pygatyphella B complex; very similar to Pygat. limbatipennis , which is not known from San Cristobal , distinguished by the median pronotal markings, the paler brown elytra, the slightly paler and much wider lateral elytral margins, and the restricted occurrence on San Cristobal . Male. 8.1–9.8 mm long; 3.4–3.9 mm wide; W/L 0.4. Colour ( Fig. 329, 338 ): Pronotum orange yellow, semitransparent with underlying fat body visible, and 2 small median dark brown areas; elytra mid-brown, lateral margins widely pale yellow and semitransparent, pale margins extending from outer edge of humerus to apical area of suture; MS and MN pale yellow; head, antennae and palpi very dark brown; venter of thorax pale orange yellow except for very dark brown tarsi and apical ¾ of tibiae; basal ventrites yellowish, V3 with dark markings in one Bweinaniawarikiapu male and two Napagiwae; V4 extensively in representatives from all locations, V4 lightly marked in Bweinaniawarikiapu (3) and Manipwena (2); V5 always very dark brown, V7 yellowish and semitransparent behind LO; basal tergites yellowish, T 4–6 marked irregularly in light brown; T 7 and 8 orange yellow and semitransparent. Pronotum ( Fig. 338 ): 1.7–2.3 mm long; 3.0– 3.7 mm wide; W/L 1.6–1.7; lateral margins with rounded convergence in posterior areas; lacking indentation near posterolateral corners; with slight irregularities along rounded posterolateral corners, which project at least as far as median posterior margin. Elytron: convex-sided; 6.0– 7.5 mm long; interstitial lines not well–defined. Head: large, moderately exposed, not able to be completely retracted into prothoracic cavity; GHW 1.9–2.4 mm ; SIW 0.2–0.3 mm ; SIW/GHW 0.1; ASD<ASW, sockets very close but not contiguous; frons not defined. Abdomen, ventrites: LO in V7 occupying half or less than half the area, not reaching sides, posterior margin not emarginated; muscle impressions clearly visible through cuticle in arched, not swollen, posterior half; posterolateral corners of V7 angulate. MPP symmetrical or asymmetrical, squarely or obliquely truncate (slightly obliquely truncate in Wugiroga 1, Bweinaniwarikiapu 1, Manipwena 3, and Napagiwae 1; obliquely truncate in remainder, very strongly in Bweinaniwarikiapu 1 and Kira Kira 1); with a broad dorsal ridge developed slightly to the left of centre. Tergites: T8 outline pattern 4 ( Fig. 440 ); as wide as long, with low rounded curved elevation in posterior area to one side of mid-line; lateral margins converge posteriorly and median posterior margin (viewed from above) narrowly rounded; anterolateral prolongations of T8 elongate, narrow, and expanded vertically; ventrally directed pieces at bases of prolongations of T8 present. Aedeagal sheath ( Figs 362, 363 ) subparallel-sided for 1/3–1/2 of its length past articulation with sheath tergite, then right side emarginate; anterior margin of tergite not emarginated; tergite with slight projection to R (as in Figs 360, 361 ). Aedeagus ( Figs 370, 371 ): L/W <3; LL/ML wide; LL diverge along most of their length dorsally, slightly shorter than ML, apices rounded and subequal in width to ML; ML bearing narrow pointed lateral teeth, apex rounded truncate; base of LL not asymmetrically produced, slightly irregularly rounded. Female. 9.0– 9.6 mm long. Coloured as for male with these exceptions: V6 with yellowish LO which is retracted narrowly across the posterior margin which is semitransparent with whitish fat body visible in irregular clumps; V7, 8 semitransparent with underlying fat body confusing the basic colour, V7 with diffuse brown markings in median and anterolateral areas in all but one Bweinaniwarikiapu female. V7 lacking anterolateral depressions; with median posterior margin shallowly emarginate, and posterolateral areas broadly rounded; V8 median posterior margin narrowly emarginate; T8 with lateral margins converging posteriorly and posterior margin rounded. Larva. Unknown. Etymology. The specific name is a noun formed by combining part of the name of the species it most resembles ( limbatipennis ) and ‘fusca’ (dark) to highlight the dark markings on the pronotum. For a discussion on the possible variability in colour and morphology in specimens of this species, Pygat . limbatipennis and Pygat. salomonis see below and Tables 8 , 9 .