<document id="43D2CEEDCB16BB346D1F13CB3DD4494A" ID-CLB-Dataset="36983" ID-DOI="10.11646/zootaxa.3702.1.1" ID-GBIF-Dataset="7cea128b-57e1-46b8-84ca-5fc9da104834" ID-ISSN="1175-5326" ID-Zenodo-Dep="284506" ID-ZooBank="84A22080-F9BB-406E-B10F-1A3A2D161C5D" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460669484449" checkinUser="plazi" docAuthor="Ripka, Geza, Laniecka, Izabella &amp; Kazmierski, Andrzej" docDate="2013" docId="038887D5FFA4EF31FF5748ACFD972F25" docLanguage="en" docName="zt03702p050.pdf" docOrigin="Zootaxa 3702 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tydeus clavimaculatus Kazmierski, sp. nov." docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="FFB1FFADFFADEF3AFFC04B1CFFF32826" masterDocTitle="On the arboreal acarofauna of Hungary: Some new and rare species of prostigmatic mites (Acari: Prostigmata: Tydeidae, Iolinidae and Stigmaeidae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="10" updateTime="1732883719967" updateUser="ExternalLinkService">
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<mods:title id="38B26F4ED4B160B30B5D4641EE3E91BC">On the arboreal acarofauna of Hungary: Some new and rare species of prostigmatic mites (Acari: Prostigmata: Tydeidae, Iolinidae and Stigmaeidae)</mods:title>
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<mods:namePart id="0F16E84928EA24F138C538A315C5E1FD">Ripka, Geza</mods:namePart>
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<mods:namePart id="5F86DDC9F319BEDEEF7D593CECD52209">Laniecka, Izabella</mods:namePart>
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<mods:date id="F18C45E3B2D0B3AB16799F667DE327B6">2013</mods:date>
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<treatment id="038887D5FFA4EF31FF5748ACFD972F25" ID-DOI="http://doi.org/10.5281/zenodo.5626212" ID-GBIF-Taxon="127665355" ID-Zenodo-Dep="5626212" LSID="urn:lsid:plazi:treatment:038887D5FFA4EF31FF5748ACFD972F25" httpUri="http://treatment.plazi.org/id/038887D5FFA4EF31FF5748ACFD972F25" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" scope_family="Tydeidae" scope_subOrder="Prostigmata">
<subSubSection id="C33B6548FFA4EF31FF5748ACFD972F25" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B9E36C3FFA4EF33FF5748ACFD542BEC" blockId="9.[151,679,944,1003]" box="[151,679,944,970]" pageId="9" pageNumber="10">
<heading id="D0D681AFFFA4EF33FF5748ACFD542BEC" bold="true" box="[151,679,944,970]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B955EAD1FFA4EF33FF5748ACFD542BEC" bold="true" box="[151,679,944,970]" pageId="9" pageNumber="10">
<taxonomicName id="4C214D40FFA4EF33FF5748ACFDB12BEC" authority="Kazmierski" authorityName="Kazmierski" box="[151,578,944,970]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="clavimaculatus" status="sp. nov.">
<emphasis id="B955EAD1FFA4EF33FF5748ACFE5A2BEC" bold="true" box="[151,425,944,970]" italics="true" pageId="9" pageNumber="10">Tydeus clavimaculatus</emphasis>
Kazmierski
</taxonomicName>
<taxonomicNameLabel id="A26657AAFFA4EF33FD8948ACFD542BEC" box="[585,679,944,970]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9E36C3FFA4EF33FF5748CEFED32BCD" blockId="9.[151,679,944,1003]" box="[151,288,978,1003]" pageId="9" pageNumber="10">
(
<figureCitation id="131A2A46FFA4EF33FF5F48CEFEEB2BCD" box="[159,280,978,1003]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStartId-0="10.[151,250,1998,2020]" captionStartId-1="11.[151,250,1393,1415]" captionStartId-2="12.[151,250,1906,1928]" captionStartId-3="13.[151,250,1509,1531]" captionTargetBox-0="[350,1254,398,1961]" captionTargetBox-1="[359,1240,210,1371]" captionTargetBox-2="[212,1375,201,1871]" captionTargetBox-3="[333,1253,193,1487]" captionTargetId-0="figure@10.[322,1264,385,1976]" captionTargetId-1="figure@11.[338,1249,193,1372]" captionTargetId-2="figure@12.[200,1387,71,1885]" captionTargetId-3="figure@13.[333,1253,193,1488]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="FIGURE 7. Tydeus clavimaculatus Kazmierski, sp. nov., holotype tritonymph: dorsal side." captionText-1="FIGURE 8. Tydeus clavimaculatus Kazmierski, sp. nov., holotype tritonymph: ventral side, genital and pseudanal region." captionText-2="FIGURE 9. Tydeus clavimaculatus Kazmierski, sp. nov., holotype tritonymph. (A) left leg I (dorsally, somewhat paraxially); (B) left leg II (dorsally, slightly paraxially); (C) cheliceral stiletto; (D) left palpus (dorso-paraxial view); (E) dorsal idiosomal setae (trichobothrium, aspidosomal ro, opisthosomal h 2)." captionText-3="FIGURE 10. Tydeus clavimaculatus Kazmierski, sp. nov., holotype tritonymph. (A) general aspect; (B) fragment of dorsal granulation (“ basketweave ”); (C) left tarsus and tibia of leg I; (D) ventral side: genital and pseudanal region." httpUri-0="https://zenodo.org/record/284513/files/figure.png" httpUri-1="https://zenodo.org/record/284514/files/figure.png" httpUri-2="https://zenodo.org/record/284515/files/figure.png" httpUri-3="https://zenodo.org/record/284516/files/figure.png" pageId="9" pageNumber="10">Figs. 7–10</figureCitation>
)
</paragraph>
<paragraph id="8B9E36C3FFA4EF33FF574F06FCEC2C69" blockId="9.[151,799,1050,1103]" pageId="9" pageNumber="10">
<taxonomicName id="4C214D40FFA4EF33FF574F06FE982C16" box="[151,363,1050,1072]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B955EAD1FFA4EF33FF574F06FF2E2C16" box="[151,221,1050,1072]" italics="true" pageId="9" pageNumber="10">Tydeus</emphasis>
cf. 
<emphasis id="B955EAD1FFA4EF33FEC34F06FE982C16" box="[259,363,1050,1072]" italics="true" pageId="9" pageNumber="10">maculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26657AAFFA4EF33FEB24F07FE5B2C16" box="[370,424,1051,1072]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
IV, Ripka &amp; Kaźmierski 1998a: 412 
<taxonomicName id="4C214D40FFA4EF33FF574F25FE982C69" box="[151,363,1081,1103]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B955EAD1FFA4EF33FF574F25FF2E2C69" box="[151,221,1081,1103]" italics="true" pageId="9" pageNumber="10">Tydeus</emphasis>
cf. 
<emphasis id="B955EAD1FFA4EF33FEC34F25FE982C69" box="[259,363,1081,1103]" italics="true" pageId="9" pageNumber="10">maculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26657AAFFA4EF33FEB24F26FE5A2C69" box="[370,425,1082,1103]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
, Çobanoglu &amp; Kaźmierski 1999: 76.
</paragraph>
<paragraph id="8B9E36C3FFA4EF33FF574F9CFE0C2D0B" blockId="9.[151,1437,1147,2012]" pageId="9" pageNumber="10">
<typeStatus id="549A8861FFA4EF33FF574F9CFF3C2CBE" box="[151,207,1152,1176]" pageId="9" pageNumber="10">Type</typeStatus>
material. Tritonymph 
<typeStatus id="549A8861FFA4EF33FE144F62FDCA2CB1" box="[468,569,1150,1175]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
, 1 tritonymph 
<typeStatus id="549A8861FFA4EF33FD254F9CFCB62CBE" box="[741,837,1152,1176]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
(slide N o 925 =AK H-295) from 
<taxonomicName id="4C214D40FFA4EF33FB0E4F62FA892CB1" box="[1230,1402,1150,1175]" class="Magnoliopsida" family="Sapindaceae" genus="Aesculus" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="pavia">
<emphasis id="B955EAD1FFA4EF33FB0E4F62FA892CB1" box="[1230,1402,1150,1175]" italics="true" pageId="9" pageNumber="10">Aesculus pavia</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C214D40FFA4EF33FF5E4FBEFE9C2C9D" box="[158,367,1186,1211]" class="Magnoliopsida" family="Hippocastanaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="family">Hippocastanaceae</taxonomicName>
), 
<collectingCountry id="F3367653FFA4EF33FE444FBFFDE22C9D" box="[388,529,1187,1211]" name="Hungary" pageId="9" pageNumber="10">HUNGARY</collectingCountry>
: Zirc (Veszprém County), 
<date id="FF9F1003FFA4EF33FC834FBEFC122C9D" box="[835,993,1186,1211]" pageId="9" pageNumber="10" value="1997-05-26">26 May 1997</date>
, coll. G. Ripka. 1 tritonymph 
<typeStatus id="549A8861FFA4EF33FAFB4FBFFA682C9D" box="[1339,1435,1187,1211]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
(slide N o 929 =AK H-297) from leaves of 
<taxonomicName id="4C214D40FFA4EF33FDA74FD6FCE62CC7" box="[615,789,1226,1249]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="campestre">
<emphasis id="B955EAD1FFA4EF33FDA74FD6FCE62CC7" box="[615,789,1226,1249]" italics="true" pageId="9" pageNumber="10">Acer campestre</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C214D40FFA4EF33FC814FD6FC4F2CC4" box="[833,956,1226,1250]" class="Magnoliopsida" family="Aceraceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="family">Aceraceae</taxonomicName>
), 
<collectingCountry id="F3367653FFA4EF33FC0E4FD6FBA82CC4" box="[974,1115,1226,1250]" name="Hungary" pageId="9" pageNumber="10">HUNGARY</collectingCountry>
: Zirc (Veszprém County), 
<date id="FF9F1003FFA4EF33FA404FD5FEF82D2F" pageId="9" pageNumber="10" value="1997-05-26">26 May 1997</date>
, coll. G. Ripka; 3 tritonymph 
<typeStatus id="549A8861FFA4EF33FD944FEDFD4C2D2F" box="[596,703,1265,1289]" pageId="9" pageNumber="10" type="paratype">paratypes</typeStatus>
(slide N o 929b =AK H-298) from leaves of 
<taxonomicName id="4C214D40FFA4EF33FB684FEDFAA62D2E" box="[1192,1365,1265,1288]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="campestre">
<emphasis id="B955EAD1FFA4EF33FB684FEDFAA62D2E" box="[1192,1365,1265,1288]" italics="true" pageId="9" pageNumber="10">Acer campestre</emphasis>
</taxonomicName>
, same locality and date, coll. G. Ripka.
</paragraph>
<paragraph id="8B9E36C3FFA4EF33FF074E27FC572D5E" blockId="9.[151,1437,1147,2012]" pageId="9" pageNumber="10">
Additional material. 2 tritonymphs (slide N o 8-330) from 
<taxonomicName id="4C214D40FFA4EF33FCB44E20FAE62D75" authority="Lipsky" authorityName="Lipsky" box="[884,1301,1339,1364]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B955EAD1FFA4EF33FCB44E20FBC22D72" box="[884,1073,1339,1364]" italics="true" pageId="9" pageNumber="10">Fagus orientalis</emphasis>
Lipsky (Fagaceae)
</taxonomicName>
, 
<collectingCountry id="F3367653FFA4EF33FAE54E20FA6B2D75" box="[1317,1432,1340,1363]" name="Turkey" pageId="9" pageNumber="10">TURKEY</collectingCountry>
: Istranca Mountains, Kirklareli, 
<date id="FF9F1003FFA4EF33FE374E43FD7C2D51" box="[503,655,1375,1399]" pageId="9" pageNumber="10" value="1992-06-14">14 June 1992</date>
, coll. Sultan Çobanoglu.
</paragraph>
<paragraph id="8B9E36C3FFA4EF33FF074E98FE3F2DBD" blockId="9.[151,1437,1147,2012]" box="[199,460,1411,1436]" pageId="9" pageNumber="10">
<typeStatus id="549A8861FFA4EF33FF074E98FF0C2DBA" box="[199,255,1412,1436]" pageId="9" pageNumber="10">Type</typeStatus>
repository. DAM.
</paragraph>
<paragraph id="8B9E36C3FFA4EF33FF074EBBFEAB2E2E" blockId="9.[151,1437,1147,2012]" pageId="9" pageNumber="10">
Etymology. The specific name is a combination of the empodial claw (present in this species) and the name of the closely related 
<taxonomicName id="4C214D40FFA4EF33FEB84ED0FCC02DC2" authority="Momen and Lundqvist" authorityName="Momen and Lundqvist" box="[376,819,1483,1508]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B955EAD1FFA4EF33FEB84ED0FDFF2DC2" box="[376,524,1483,1508]" italics="true" pageId="9" pageNumber="10">T. maculatus</emphasis>
(Momen and Lundqvist)
</taxonomicName>
(in which the empodial claw is absent) (Momen &amp; Lundqvist 1996).
</paragraph>
<paragraph id="8B9E36C3FFA4EF33FF074D0FFA9F2E76" blockId="9.[151,1437,1147,2012]" pageId="9" pageNumber="10">
Relation to host. Underside of leaves of 
<taxonomicName id="4C214D40FFA4EF33FD774D0FFC942E0D" box="[695,871,1555,1580]" class="Magnoliopsida" family="Sapindaceae" genus="Aesculus" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="pavia">
<emphasis id="B955EAD1FFA4EF33FD774D0FFC942E0D" box="[695,871,1555,1580]" italics="true" pageId="9" pageNumber="10">Aesculus pavia</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C214D40FFA4EF33FCBC4D0FFC062E0A" box="[892,1013,1555,1580]" class="Insecta" family="Aphididae" kingdom="Animalia" order="Aphidomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Aphididae</taxonomicName>
: 
<taxonomicName id="4C214D40FFA4EF33FBC54D0FFA6B2E0A" authority="Fernie" authorityName="Fernie" box="[1029,1432,1555,1580]" class="Insecta" family="Aphididae" genus="Periphyllus" kingdom="Animalia" order="Aphidomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="testudinaceus">
<emphasis id="B955EAD1FFA4EF33FBC54D0FFADE2E0A" box="[1029,1325,1555,1580]" italics="true" pageId="9" pageNumber="10">Periphyllus testudinaceus</emphasis>
(Fernie)
</taxonomicName>
; 
<taxonomicName id="4C214D40FFA4EF33FF574D2BFED82E76" box="[151,299,1591,1616]" class="Arachnida" family="Phytoseiidae" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Phytoseiidae</taxonomicName>
: 
<taxonomicName id="4C214D40FFA4EF33FEF44D24FD572E76" authority="Oudemans" authorityName="Oudemans" box="[308,676,1591,1616]" class="Arachnida" family="Phytoseiidae" genus="Euseius" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="finlandicus">
<emphasis id="B955EAD1FFA4EF33FEF44D24FDFC2E76" box="[308,527,1591,1616]" italics="true" pageId="9" pageNumber="10">Euseius finlandicus</emphasis>
(Oudemans)
</taxonomicName>
; 
<taxonomicName id="4C214D40FFA4EF33FD6F4D2BFCE92E76" box="[687,794,1591,1616]" class="Arachnida" family="Tydeidae" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Tydeidae</taxonomicName>
: 
<taxonomicName id="4C214D40FFA4EF33FCE44D2BFB9F2E76" authority="Castagnoli" authorityName="Castagnoli" box="[804,1132,1591,1616]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="calabrus">
<emphasis id="B955EAD1FFA4EF33FCE44D2BFC292E76" box="[804,986,1591,1616]" italics="true" pageId="9" pageNumber="10">Tydeus calabrus</emphasis>
(Castagnoli)
</taxonomicName>
[lower side of leaves].
</paragraph>
<paragraph id="8B9E36C3FFA4EF33FF074D40FD4D2E9A" blockId="9.[151,1437,1147,2012]" pageId="9" pageNumber="10">
<emphasis id="B955EAD1FFA4EF33FF074D40FE7A2E55" box="[199,393,1628,1651]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C214D40FFA4EF33FF074D40FE762E55" box="[199,389,1628,1651]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="campestre">Acer campestre</taxonomicName>
.
</emphasis>
<taxonomicName id="4C214D40FFA4EF33FE5C4D47FDDA2E52" box="[412,553,1627,1652]" class="Arachnida" family="Eriophyidae" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Eriophyidae</taxonomicName>
: 
<taxonomicName id="4C214D40FFA4EF33FD814D40FC6A2E52" authority="Nalepa" authorityName="Nalepa" box="[577,921,1627,1652]" class="Arachnida" family="Eriophyidae" genus="Aceria" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cephalonea">
<emphasis id="B955EAD1FFA4EF33FD814D40FCD32E52" box="[577,800,1627,1652]" italics="true" pageId="9" pageNumber="10">Aceria cephalonea</emphasis>
(Nalepa)
</taxonomicName>
; 
<taxonomicName id="4C214D40FFA4EF33FC714D47FBEF2E52" box="[945,1052,1627,1652]" class="Arachnida" family="Tydeidae" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Tydeidae</taxonomicName>
: 
<taxonomicName id="4C214D40FFA4EF33FBF34D47FA6B2E52" authority="Castagnoli" authorityName="Castagnoli" box="[1075,1432,1627,1652]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="calabrus">
<emphasis id="B955EAD1FFA4EF33FBF34D47FB052E52" box="[1075,1270,1627,1652]" italics="true" pageId="9" pageNumber="10">Tydeus calabrus</emphasis>
(Castagnoli)
</taxonomicName>
; 
<taxonomicName id="4C214D40FFA4EF33FF574D63FEDC2EBE" box="[151,303,1663,1688]" class="Arachnida" family="Meyerellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Meyerellidae</taxonomicName>
: 
<taxonomicName id="4C214D40FFA4EF33FE874D63FCDC2EBE" authority="Oudemans" authorityName="Oudemans" box="[327,815,1663,1688]" class="Arachnida" family="Tydeidae" genus="Triophtydeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="triophthalmus">
<emphasis id="B955EAD1FFA4EF33FE874D63FD7F2EBE" box="[327,652,1663,1688]" italics="true" pageId="9" pageNumber="10">Triophtydeus triophthalmus</emphasis>
(Oudemans)
</taxonomicName>
; 
<taxonomicName id="4C214D40FFA4EF33FC894D63FBDA2EBE" box="[841,1065,1663,1688]" class="Arachnida" family="Winterschmidtiidae" kingdom="Animalia" order="Astigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Winterschmidtiidae</taxonomicName>
: 
<taxonomicName id="4C214D40FFA4EF33FB824D63FED12E9A" authority="Oudemans" authorityName="Oudemans" class="Arachnida" family="Winterschmidtiidae" genus="Czenspinskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="trasversostriata">
<emphasis id="B955EAD1FFA4EF33FB824D63FA6F2EB1" box="[1090,1436,1663,1688]" italics="true" pageId="9" pageNumber="10">Czenspinskia trasversostriata</emphasis>
(Oudemans)
</taxonomicName>
[underside of leaves of field maple].
</paragraph>
<paragraph id="8B9E36C3FFA4EF33FF074DDBFBD22F6D" blockId="9.[151,1437,1147,2012]" pageId="9" pageNumber="10">
Differential diagnosis. The new species is similar to 
<taxonomicName id="4C214D40FFA4EF33FCB34DDBFBB42EC6" box="[883,1095,1735,1760]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B955EAD1FFA4EF33FCB34DDBFBB42EC6" box="[883,1095,1735,1760]" italics="true" pageId="9" pageNumber="10">Tydeus maculatus</emphasis>
</taxonomicName>
, mainly because of dorsal ornamentation (”Tydulosus” 
<typeStatus id="549A8861FFA4EF33FE294DF0FDEF2F22" box="[489,540,1772,1796]" pageId="9" pageNumber="10">type</typeStatus>
, or “basket weave”—see Kaźmierski [1998b]). However, it differs by two features: [1] the presence of the empodial claws (absent in 
<taxonomicName id="4C214D40FFA4EF33FC894C0CFC282F0E" box="[841,987,1807,1832]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B955EAD1FFA4EF33FC894C0CFC282F0E" box="[841,987,1807,1832]" italics="true" pageId="9" pageNumber="10">T. maculatus</emphasis>
</taxonomicName>
), and [2] the shape of the tip of tarsal eupathidium 
<emphasis id="B955EAD1FFA4EF33FEEA4C2FFEB02F6A" box="[298,323,1843,1868]" italics="true" pageId="9" pageNumber="10">pζ</emphasis>
, which is forked distally (instead of “T”-shaped in 
<taxonomicName id="4C214D40FFA4EF33FC434C28FBE22F6A" box="[899,1041,1843,1868]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B955EAD1FFA4EF33FC434C28FBE22F6A" box="[899,1041,1843,1868]" italics="true" pageId="9" pageNumber="10">T. maculatus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B9E36C3FFA4EF33FF074C4BFA6E2FFA" blockId="9.[151,1437,1147,2012]" pageId="9" pageNumber="10">
Description. TRITONYMPH (
<typeStatus id="549A8861FFA4EF33FDE24C4BFD782F56" box="[546,651,1879,1904]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
). Organotaxy of gnathosoma, idiosoma and appendages—typical for genus (Kaźmierski 1989b). 
<emphasis id="B955EAD1FFA4EF33FE104C67FDCB2FB2" box="[464,568,1915,1940]" italics="true" pageId="9" pageNumber="10">Idiosoma</emphasis>
. Body oval, yellowish. Length 245, width 185. 
<typeStatus id="549A8861FFA4EF33FB964C60FB4B2FB2" box="[1110,1208,1916,1940]" pageId="9" pageNumber="10" type="paratype">Paratype</typeStatus>
tritonymphs: length 240–245 / width 180–185. Dorsally, laterally and on the circumference of ventral side covered with granulations, “basket weave”. Striae between setae 
<emphasis id="B955EAD1FFA4EF33FD904CDFFD9F2FFA" box="[592,620,1987,2012]" italics="true" pageId="9" pageNumber="10">d1</emphasis>
transverse. On aspidosoma striae are longitudinal. Eyes not observed.
</paragraph>
<paragraph id="8B9E36C3FFA7EF31FF574B8BFBA12E2F" blockId="10.[151,1436,151,320]" lastBlockId="11.[151,1437,1454,1795]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Bothridial setae 
<emphasis id="B955EAD1FFA7EF30FE954B8BFE772896" box="[341,388,151,176]" italics="true" pageId="10" pageNumber="11">(bo)</emphasis>
whip-like, rather flabby than stout and longer than the other dorsal body setae. Latter are serrated and representing 2 different kinds: (1) narrowly lanceolate and pointed distally: 
<emphasis id="B955EAD1FFA7EF30FB424BA1FAE328F2" box="[1154,1296,188,213]" italics="true" pageId="10" pageNumber="11">ro, la, ex, c1</emphasis>
, 
<emphasis id="B955EAD1FFA7EF30FADF4BA0FACA28F2" box="[1311,1337,188,212]" italics="true" pageId="10" pageNumber="11">c2</emphasis>
, 
<emphasis id="B955EAD1FFA7EF30FA884BA0FA9628F3" box="[1352,1381,188,213]" italics="true" pageId="10" pageNumber="11">d1</emphasis>
, and (2) flat, spatulate: 
<emphasis id="B955EAD1FFA7EF30FEAA4BFCFE7728DE" box="[362,388,224,248]" italics="true" pageId="10" pageNumber="11">e1</emphasis>
, 
<emphasis id="B955EAD1FFA7EF30FE544BFCFE5A28DE" box="[404,425,224,248]" italics="true" pageId="10" pageNumber="11">f1</emphasis>
, 
<emphasis id="B955EAD1FFA7EF30FE7A4BFCFE3C28DE" box="[442,463,224,248]" italics="true" pageId="10" pageNumber="11">f2</emphasis>
, 
<emphasis id="B955EAD1FFA7EF30FE1F4BC3FE0828DE" box="[479,507,223,248]" italics="true" pageId="10" pageNumber="11">h1</emphasis>
, 
<emphasis id="B955EAD1FFA7EF30FDCB4BC3FDDB28DE" box="[523,552,223,248]" italics="true" pageId="10" pageNumber="11">h2</emphasis>
and 
<emphasis id="B955EAD1FFA7EF30FDA14BFCFD7B28DE" box="[609,648,224,248]" italics="true" pageId="10" pageNumber="11">ps1</emphasis>
. Lengths as follows: 
<emphasis id="B955EAD1FFA7EF30FC404BC3FC6F28DE" box="[896,924,223,248]" italics="true" pageId="10" pageNumber="11">bo</emphasis>
37, 
<emphasis id="B955EAD1FFA7EF30FC114BFDFC1A28DE" box="[977,1001,225,248]" italics="true" pageId="10" pageNumber="11">ro</emphasis>
15, 
<emphasis id="B955EAD1FFA7EF30FBDD4BC3FBC028DE" box="[1053,1075,223,248]" italics="true" pageId="10" pageNumber="11">la</emphasis>
18, 
<emphasis id="B955EAD1FFA7EF30FBA84BFDFB7228DE" box="[1128,1153,225,248]" italics="true" pageId="10" pageNumber="11">ex</emphasis>
20, 
<emphasis id="B955EAD1FFA7EF30FB754BFCFB3C28DE" box="[1205,1231,224,248]" italics="true" pageId="10" pageNumber="11">c1</emphasis>
18, 
<emphasis id="B955EAD1FFA7EF30FAC54BFCFAEC28DE" box="[1285,1311,224,248]" italics="true" pageId="10" pageNumber="11">c2</emphasis>
20, 
<emphasis id="B955EAD1FFA7EF30FA944BC3FA9C28DE" box="[1364,1391,223,248]" italics="true" pageId="10" pageNumber="11">d1</emphasis>
18, remaining setae (
<emphasis id="B955EAD1FFA7EF30FE984A18FE80293A" box="[344,371,260,284]" italics="true" pageId="10" pageNumber="11">e1</emphasis>
– 
<emphasis id="B955EAD1FFA7EF30FE404A18FE54293A" box="[384,423,260,284]" italics="true" pageId="10" pageNumber="11">ps1</emphasis>
, spatulate)—15. Setae 
<emphasis id="B955EAD1FFA7EF30FD744A18FD29293A" box="[692,730,260,284]" italics="true" pageId="10" pageNumber="11">ps2</emphasis>
acute 12. Distances: 
<emphasis id="B955EAD1FFA7EF30FC0A4A18FBFE293A" box="[970,1037,260,284]" italics="true" pageId="10" pageNumber="11">c1–c1</emphasis>
43, 
<emphasis id="B955EAD1FFA7EF30FBFE4A18FB77293B" box="[1086,1156,260,285]" italics="true" pageId="10" pageNumber="11">d1–d1</emphasis>
32, 
<emphasis id="B955EAD1FFA7EF30FB764A18FB0A293A" box="[1206,1273,260,284]" italics="true" pageId="10" pageNumber="11">e1–e1</emphasis>
105, 
<emphasis id="B955EAD1FFA7EF30FAF84A18FA81293A" box="[1336,1394,260,284]" italics="true" pageId="10" pageNumber="11">f1–f1</emphasis>
48, 
<emphasis id="B955EAD1FFA7EF30FF574A3BFF2E2966" box="[151,221,295,320]" italics="true" pageId="10" pageNumber="11">h1–h1</emphasis>
48, 
<emphasis id="B955EAD1FFA7EF30FECC4A34FE9B2966" box="[268,360,296,320]" italics="true" pageId="10" pageNumber="11">ps1–ps1</emphasis>
14, 
<emphasis id="B955EAD1FFA7EF30FE584A3BFE2B2966" box="[408,472,295,320]" italics="true" pageId="10" pageNumber="11">f1–h1</emphasis>
27. Lyrifissures invisible, probably hidden between granules of ornamentation. Coxal organ (
<emphasis id="B955EAD1FFA6EF31FF294EB3FEF02DE0" box="[233,259,1455,1478]" italics="true" pageId="11" pageNumber="12">cg</emphasis>
) oval. Venter striated medially. Striae longitudinal in metasternal region. Genital region with 2 genital pores flanked by 4 pairs of genital and aggenital setae: genital setal formula: (4-4). Paraproctal lips prominent and strongly striated. Setae 
<emphasis id="B955EAD1FFA6EF31FE5D4EEDFE372E2F" box="[413,452,1521,1545]" italics="true" pageId="11" pageNumber="12">ps2</emphasis>
laterally of and close to lips. Epimeral formula: (3-1-4-2).
</paragraph>
<caption id="DF5E664BFFA7EF30FF574CD2FBC72FC2" box="[151,1076,1998,2020]" httpUri="https://zenodo.org/record/284513/files/figure.png" pageId="10" pageNumber="11" targetBox="[350,1254,398,1961]" targetPageId="10">
<paragraph id="8B9E36C3FFA7EF30FF574CD2FBC72FC2" blockId="10.[151,1076,1998,2020]" box="[151,1076,1998,2020]" pageId="10" pageNumber="11">
<emphasis id="B955EAD1FFA7EF30FF574CD2FEE02FC2" bold="true" box="[151,275,1998,2020]" pageId="10" pageNumber="11">FIGURE 7.</emphasis>
<taxonomicName id="4C214D40FFA7EF30FED94CD2FD8F2FC2" authority="Kazmierski" authorityName="Kazmierski" box="[281,636,1998,2020]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="clavimaculatus" status="sp. nov.">
<emphasis id="B955EAD1FFA7EF30FED94CD2FE0D2FC2" box="[281,510,1998,2020]" italics="true" pageId="10" pageNumber="11">Tydeus clavimaculatus</emphasis>
Kazmierski
</taxonomicName>
, 
<emphasis id="B955EAD1FFA7EF30FD464CD2FD252FC2" bold="true" box="[646,726,1998,2020]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A26657AAFFA7EF30FD464CD2FD252FC2" box="[646,726,1998,2020]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype tritonymph: dorsal side.
</paragraph>
</caption>
<caption id="DF5E664BFFA6EF31FF574E6DFA9F2DA1" box="[151,1388,1393,1415]" httpUri="https://zenodo.org/record/284514/files/figure.png" pageId="11" pageNumber="12" targetBox="[359,1240,210,1371]" targetPageId="11">
<paragraph id="8B9E36C3FFA6EF31FF574E6DFA9F2DA1" blockId="11.[151,1388,1393,1415]" box="[151,1388,1393,1415]" pageId="11" pageNumber="12">
<emphasis id="B955EAD1FFA6EF31FF574E6DFEE02DA1" bold="true" box="[151,275,1393,1415]" pageId="11" pageNumber="12">FIGURE 8.</emphasis>
<taxonomicName id="4C214D40FFA6EF31FED94E6DFD8F2DA1" authority="Kazmierski" authorityName="Kazmierski" box="[281,636,1393,1415]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="clavimaculatus" status="sp. nov.">
<emphasis id="B955EAD1FFA6EF31FED94E6DFE0D2DA1" box="[281,510,1393,1415]" italics="true" pageId="11" pageNumber="12">Tydeus clavimaculatus</emphasis>
Kazmierski
</taxonomicName>
, 
<emphasis id="B955EAD1FFA6EF31FD464E6DFD252DA1" bold="true" box="[646,726,1393,1415]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="A26657AAFFA6EF31FD464E6DFD252DA1" box="[646,726,1393,1415]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype tritonymph: ventral side, genital and pseudanal region.
</paragraph>
</caption>
<paragraph id="8B9E36C3FFA6EF31FF074D0EFDE42EB1" blockId="11.[151,1437,1454,1795]" pageId="11" pageNumber="12">
Gnathosoma. Visible from above. Cheliceral stilettos similar in length to palpal tarsus (only slightly longer). Eupathidium 
<emphasis id="B955EAD1FFA6EF31FEF54D2AFE922E69" box="[309,353,1590,1615]" italics="true" pageId="11" pageNumber="12">(pζ)</emphasis>
insignificantly bent, forked distally. Seta 
<emphasis id="B955EAD1FFA6EF31FC9C4D2AFC992E69" box="[860,874,1590,1615]" italics="true" pageId="11" pageNumber="12">d</emphasis>
forked distally. Minute seta 
<emphasis id="B955EAD1FFA6EF31FB0B4D2AFB142E69" box="[1227,1255,1590,1615]" italics="true" pageId="11" pageNumber="12">ba</emphasis>
relatively well developed. Palpal chaetotaxy: 6+ 
<emphasis id="B955EAD1FFA6EF31FDD04D47FDD72E54" box="[528,548,1627,1650]" italics="true" pageId="11" pageNumber="12">ω</emphasis>
–2 –2. Measurements: stiletto 15.5, palpal femuro-genu 20/9, 
<emphasis id="B955EAD1FFA6EF31FB304D46FAF52E55" box="[1264,1286,1626,1651]" italics="true" pageId="11" pageNumber="12">df</emphasis>
12, 
<emphasis id="B955EAD1FFA6EF31FAFA4D46FAA52E55" box="[1338,1366,1626,1651]" italics="true" pageId="11" pageNumber="12">dg</emphasis>
13, 
<emphasis id="B955EAD1FFA6EF31FA4B4D47FA6F2E54" box="[1419,1436,1627,1650]" italics="true" pageId="11" pageNumber="12">t’</emphasis>
15, 
<emphasis id="B955EAD1FFA6EF31FF014D63FF2B2EB0" box="[193,216,1663,1686]" italics="true" pageId="11" pageNumber="12">t’’</emphasis>
8, palpal tarsus 14/5, 
<emphasis id="B955EAD1FFA6EF31FE0E4D62FE082EB1" box="[462,507,1662,1687]" italics="true" pageId="11" pageNumber="12">(pζ)</emphasis>
5.
</paragraph>
<paragraph id="8B9E36C3FFA6EF31FF074DBFFD972F25" blockId="11.[151,1437,1454,1795]" pageId="11" pageNumber="12">
Legs. Setation: I (8+ 
<emphasis id="B955EAD1FFA6EF31FE724DBFFE352E9C" box="[434,454,1699,1722]" italics="true" pageId="11" pageNumber="12">ω</emphasis>
-3+ 
<emphasis id="B955EAD1FFA6EF31FE2D4DBEFDFA2E9D" box="[493,521,1698,1723]" italics="true" pageId="11" pageNumber="12">k’’</emphasis>
-3-3-1), II (6+ 
<emphasis id="B955EAD1FFA6EF31FD6A4DBFFD4D2E9C" box="[682,702,1699,1722]" italics="true" pageId="11" pageNumber="12">ω</emphasis>
-2-2-2-0), III (5-2-1-1-1), IV (5-2-1-1-0). Solenidiotaxy: 2 (
<emphasis id="B955EAD1FFA6EF31FAA64DBFFA702E9C" box="[1382,1411,1699,1722]" italics="true" pageId="11" pageNumber="12">ωI</emphasis>
+ 
<emphasis id="B955EAD1FFA6EF31FF574DDBFF4E2EF8" box="[151,189,1735,1758]" italics="true" pageId="11" pageNumber="12">ωII</emphasis>
). Tarsus+apotele I: length 33, width 12, height 13. Length of 
<emphasis id="B955EAD1FFA6EF31FCBF4DDBFC6F2EF8" box="[895,924,1735,1758]" italics="true" pageId="11" pageNumber="12">ωI</emphasis>
: 4, 
<emphasis id="B955EAD1FFA6EF31FC0A4DDBFC102EF9" box="[970,995,1735,1759]" italics="true" pageId="11" pageNumber="12">ft’</emphasis>
12, 
<emphasis id="B955EAD1FFA6EF31FBD64DDAFBB32EF9" box="[1046,1088,1734,1759]" italics="true" pageId="11" pageNumber="12">ft’’ζ</emphasis>
18, 
<emphasis id="B955EAD1FFA6EF31FBB54DDAFB622EF9" box="[1141,1169,1734,1759]" italics="true" pageId="11" pageNumber="12">k’’</emphasis>
4 (forked distally), 
<emphasis id="B955EAD1FFA6EF31FAB54DDBFA6F2EF8" box="[1397,1436,1735,1758]" italics="true" pageId="11" pageNumber="12">ωII</emphasis>
1.5. Empodial hooks 
<emphasis id="B955EAD1FFA6EF31FE454DF7FE482F24" box="[389,443,1771,1794]" italics="true" pageId="11" pageNumber="12">(om)</emphasis>
present, small.
</paragraph>
</subSubSection>
</treatment>
</document>