<document id="5932E7D25927388EB14856C20C55E15A" ID-DOI="10.37828/em.2025.87.1" ID-ISSN="2336-9744" ID-Zenodo-Dep="16959124" ID-ZooBank="7E42C152-705D-4084-A80C-89A946B68738" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1756269195110" checkinUser="felipe" docAuthor="Volynkin, Anton V. &amp; Černý, Karel" docDate="2025" docId="038875429579182EFF5FF9B3E2F6F289" docLanguage="en" docName="EcolMontenegrina.87.1-45.pdf" docOrigin="Ecologica Montenegrina 87" docSource="https://doi.org/10.37828/em.2025.87.1" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.3:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="3" docTitle="Hollowayosia cardamoma Volynkin &amp; Černý, 2025, sp. n." docType="treatment" docVersion="2" lastPageNumber="17" masterDocId="FFB10D3A9576183EFFE3FFD9E644F16C" masterDocTitle="On the taxonomy of the genera Katha Moore, Tarika Moore, Cernyia Bucsek and Churingosia Volynkin &amp; Černý, stat. n. with the description of a new genus and thirteen new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina)" masterLastPageNumber="45" masterPageNumber="1" pageNumber="16" updateTime="1756275904593" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1FC382AFB62556E71F13B3D4BE9B6139">On the taxonomy of the genera Katha Moore, Tarika Moore, Cernyia Bucsek and Churingosia Volynkin &amp; Černý, stat. n. with the description of a new genus and thirteen new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina)</mods:title>
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<mods:namePart id="CC1CFFA4A0DC56D295914C3FD19A9EC6">Volynkin, Anton V.</mods:namePart>
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<mods:affiliation id="CE7CE32D2FDC28F3596DD2CC7BB26044">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia. E-mail: monstruncusarctia @ gmail. com; https: // orcid. org / 0000 - 0001 - 9447 - 4925</mods:affiliation>
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<mods:namePart id="0EA1CB367D93AB40946283F16442853D">Černý, Karel</mods:namePart>
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<mods:affiliation id="B848C379B4E132A93EAE13FED07C3998">Tiergartenstrasse 27, A- 6020 Innsbruck, Austria. E-mail: natura. cerny @ aon. at; https: // orcid. org / 0000 - 0001 - 7207 - 2255</mods:affiliation>
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<taxonomicName id="4C21BFD795791831FF5FF9B3E7B3F7E9" ID-CoL="KZ356" authorityName="Volynkin &amp; Černý" authorityYear="2025" box="[188,503,1642,1669]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cardamoma" status="sp. nov.">
<emphasis id="B955184695791831FF5FF9B3E7B3F7E9" bold="true" box="[188,503,1642,1669]" italics="true" pageId="15" pageNumber="16">Hollowayosia cardamoma</emphasis>
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https://zoobank.org/ 
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urn:lsid:zoobank.org:act: 
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<paragraph id="8B9EC45495791831FF5FF972E78DF7A9" blockId="15.[188,1178,1642,1733]" box="[188,457,1707,1733]" pageId="15" pageNumber="16">
(
<figureCitation id="131AD8D195791831FF27F972E708F7A9" box="[196,332,1707,1733]" captionStart="Figures 57–68" captionStartId="25.[188,274,1907,1931]" captionTargetBox="[192,1397,188,1856]" captionTargetId="figure-68@25.[188,1399,188,1898]" captionTargetPageId="25" captionText="Figures 57–68. Adults of Cernyia (57, 58), Hollowayosia gen. n. (59–62), Churingosia (63–66) and Manulea (67, 68) spp. Depositories of the specimens: 57, 67 and 68 in MfN; 58, 61 and 66 in MWM/ZSM (61: ex CKC); 59, 60 and 62 in CKC; 63–65 in NHMUK." figureDoi="http://doi.org/10.5281/zenodo.16959142" httpUri="https://zenodo.org/record/16959142/files/figure.png" pageId="15" pageNumber="16">Figs 61, 62</figureCitation>
, 
<figureCitation id="131AD8D195791831FEBBF972E7C3F7A9" box="[344,391,1707,1733]" captionStart="Figures 116–119" captionStartId="38.[188,274,1947,1971]" captionTargetBox="[218,1334,192,1913]" captionTargetId="figure-59@38.[202,1385,188,1928]" captionTargetPageId="38" captionText="Figures 116–119. Male genitalia of Hollowayosia gen. n. (116, 117), Churingosia nigripes (118) and Manulea palliatella (119). Depositories of the specimens dissected: 116 in CKC; 117 in MWM/ZSM; 118 in OUMNH; 119 in MfN." figureDoi="http://doi.org/10.5281/zenodo.16959188" httpUri="https://zenodo.org/record/16959188/files/figure.png" pageId="15" pageNumber="16">117</figureCitation>
, 
<figureCitation id="131AD8D195791831FE70F972E784F7A9" box="[403,448,1707,1733]" captionStart="Figures 147–152" captionStartId="44.[188,274,1944,1968]" captionTargetBox="[190,1369,207,1915]" captionTargetId="figure-65@44.[188,1369,188,1925]" captionTargetPageId="44" captionText="Figures 147–152. Female genitalia of Hollowayosia gen. n. (147, 148), Manulea palliatella (149) and Churingosia spp. (150–152). Depositories of the specimens dissected: 147 and 148 in CKC; 149 in MfN; 150 and 152 in NHMUK; 151 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.16959210" httpUri="https://zenodo.org/record/16959210/files/figure.png" pageId="15" pageNumber="16">148</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33B97DF95791831FF5FF936E52CF6FC" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8B9EC45495791831FF5FF936E419F620" blockId="15.[188,1400,1774,2037]" pageId="15" pageNumber="16">
<emphasis id="B955184695791831FF5FF936E72BF664" bold="true" box="[188,367,1774,1801]" pageId="15" pageNumber="16">Type material</emphasis>
. 
<materialsCitation id="3B49CE0995791831FE9CF936E419F620" collectingDate="2010-10-30" collectionCode="MWM, ZSM, CKC" collectorName="K. Cerny &amp; A. Volynkin" country="Thailand" elevation="150" latitude="12.802834" location="Chanthaburi" longLatPrecision="13" longitude="102.29217" pageId="15" pageNumber="16" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chanthaburi" typeStatus="holotype">
<emphasis id="B955184695791831FE9CF936E7B6F665" bold="true" box="[383,498,1775,1801]" pageId="15" pageNumber="16">
<typeStatus id="549A7AF695791831FE9CF936E7B6F665" box="[383,498,1775,1801]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="131AD8D195791831FDE0F936E424F664" box="[515,608,1774,1801]" captionStart="Figures 57–68" captionStartId="25.[188,274,1907,1931]" captionTargetBox="[192,1397,188,1856]" captionTargetId="figure-68@25.[188,1399,188,1898]" captionTargetPageId="25" captionText="Figures 57–68. Adults of Cernyia (57, 58), Hollowayosia gen. n. (59–62), Churingosia (63–66) and Manulea (67, 68) spp. Depositories of the specimens: 57, 67 and 68 in MfN; 58, 61 and 66 in MWM/ZSM (61: ex CKC); 59, 60 and 62 in CKC; 63–65 in NHMUK." figureDoi="http://doi.org/10.5281/zenodo.16959142" httpUri="https://zenodo.org/record/16959142/files/figure.png" pageId="15" pageNumber="16">Figs 61</figureCitation>
, 
<figureCitation id="131AD8D195791831FD8DF936E4DAF665" box="[622,670,1775,1801]" captionStart="Figures 116–119" captionStartId="38.[188,274,1947,1971]" captionTargetBox="[218,1334,192,1913]" captionTargetId="figure-59@38.[202,1385,188,1928]" captionTargetPageId="38" captionText="Figures 116–119. Male genitalia of Hollowayosia gen. n. (116, 117), Churingosia nigripes (118) and Manulea palliatella (119). Depositories of the specimens dissected: 116 in CKC; 117 in MWM/ZSM; 118 in OUMNH; 119 in MfN." figureDoi="http://doi.org/10.5281/zenodo.16959188" httpUri="https://zenodo.org/record/16959188/files/figure.png" pageId="15" pageNumber="16">117</figureCitation>
): male, “SE 
<collectingCountry id="F33684C495791831FCD4F936E5E4F665" box="[823,928,1775,1801]" name="Thailand" pageId="15" pageNumber="16">Thailand</collectingCountry>
| Prov. 
<collectingRegion id="49E50AB695791831FBE3F936E2D0F665" box="[1024,1172,1775,1801]" country="Thailand" name="Chanthaburi" pageId="15" pageNumber="16">Chanthaburi</collectingRegion>
, 
<quantity id="4CD969B195791831FB42F937E2B4F665" box="[1185,1264,1774,1801]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="m" value="150.0">
<elevation id="000C236795791831FB42F937E2B4F665" box="[1185,1264,1774,1801]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="m" value="150.0">150 m</elevation>
</quantity>
, | 
<quantity id="4CD969B195791831FAEEF936E31CF665" box="[1293,1368,1775,1801]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="15" pageNumber="16" unit="km" value="33.0">33 km</quantity>
N 
<collectingRegion id="49E50AB695791831FF5FF8C9E709F646" box="[188,333,1808,1834]" country="Thailand" name="Chanthaburi" pageId="15" pageNumber="16">Chanthaburi</collectingRegion>
| 
<geoCoordinate id="EE15A29395791831FE81F8C9E7AEF646" box="[354,490,1808,1834]" degrees="12" direction="north" minutes="48,17" orientation="latitude" pageId="15" pageNumber="16" precision="9" value="12.802834">12°48,17'N</geoCoordinate>
, 
<geoCoordinate id="EE15A29395791831FE15F8C9E4CEF647" box="[502,650,1808,1835]" degrees="102" direction="east" minutes="17,53" orientation="longitude" pageId="15" pageNumber="16" precision="9" value="102.29217">102°17,53'E</geoCoordinate>
, | 
<specimenCount id="9D270FDD95791831FD40F8C8E4FDF647" box="[675,697,1809,1835]" count="1" pageId="15" pageNumber="16" type="female">♀</specimenCount>
<date id="FF9FE29495791831FD22F8C9E508F646" box="[705,844,1808,1835]" pageId="15" pageNumber="16" value="2010-10-30">
<collectingDate id="EFDB1B7C95791831FD22F8C9E508F646" box="[705,844,1808,1835]" pageId="15" pageNumber="16" value="2010-10-30">30. X. 2010</collectingDate>
</date>
ex L.[arva] 
<collectorName id="26D4A18295791831FC3EF8C8E20FF642" box="[989,1099,1809,1838]" pageId="15" pageNumber="16">K. Černý</collectorName>
” / “Slide | AV8225 
<specimenCount id="9D270FDD95791831FAD3F8C8E302F647" box="[1328,1350,1809,1835]" count="1" pageId="15" pageNumber="16" type="male">♂</specimenCount>
| 
<collectorName id="26D4A18295791831FAB8F8C9E76AF620" pageId="15" pageNumber="16">A. Volynkin</collectorName>
” (
<collectionCode id="ED305C9195791831FEA4F8EBE7DEF620" box="[327,410,1842,1868]" pageId="15" pageNumber="16">MWM</collectionCode>
/ 
<collectionCode id="ED305C9195791831FE40F8EBE7A5F620" box="[419,481,1842,1868]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="15" pageNumber="16" type="Museum">ZSM</collectionCode>
, ex 
<collectionCode id="ED305C9195791831FDECF8EBE414F620" box="[527,592,1842,1868]" pageId="15" pageNumber="16">CKC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9EC45495791831FEF9F88DE52CF6FC" blockId="15.[188,1400,1774,2037]" pageId="15" pageNumber="16">
<materialsCitation id="3B49CE0995791831FEF9F88DE566F602" box="[282,802,1875,1902]" country="Thailand" location="THAILAND" pageId="15" pageNumber="16" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B955184695791831FEF9F88DE7DCF602" bold="true" box="[282,408,1876,1902]" pageId="15" pageNumber="16">
<typeStatus id="549A7AF695791831FEF9F88DE7DCF602" box="[282,408,1876,1902]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
. 
<emphasis id="B955184695791831FE45F88AE400F601" bold="true" box="[422,580,1875,1901]" pageId="15" pageNumber="16">
<collectingCountry id="F33684C495791831FE45F88AE400F601" box="[422,580,1875,1901]" name="Thailand" pageId="15" pageNumber="16">THAILAND</collectingCountry>
</emphasis>
: 
<specimenCount id="9D270FDD95791831FDB1F88DE4E7F602" box="[594,675,1876,1902]" count="1" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
, 
<specimenCount id="9D270FDD95791831FD4DF88DE566F602" box="[686,802,1876,1902]" count="2" pageId="15" pageNumber="16" type="female">2 females</specimenCount>
</materialsCitation>
, 
<materialsCitation id="3B49CE0995791831FCCFF88DE79BF6E3" country="Thailand" location="Nos." pageId="15" pageNumber="16" specimenCount="1" typeStatus="paratype">
same data as in 
<typeStatus id="549A7AF695791831FC02F88DE20CF602" box="[993,1096,1876,1902]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
but not reared from larva, gen. slide 
<location id="8EFE928F95791831FED7F8AFE72BF6FC" LSID="urn:lsid:plazi:treatment:038875429579182EFF5FF9B3E2F6F289:8EFE928F95791831FED7F8AFE72BF6FC" box="[308,367,1910,1936]" country="Thailand" name="Nos." pageId="15" pageNumber="16">Nos.</location>
: AV8226
</materialsCitation>
<specimenCount id="9D270FDD95791831FE3CF8AFE7B1F6FC" box="[479,501,1910,1936]" count="1" pageId="15" pageNumber="16" type="female">♀</specimenCount>
, AV9238 
<specimenCount id="9D270FDD95791831FD8AF8AFE43BF6FC" box="[617,639,1910,1936]" count="1" pageId="15" pageNumber="16" type="male">♂</specimenCount>
, AV9239 
<materialsCitation id="3B49CE0995791831FD10F8AFE520F6FC" box="[755,868,1910,1936]" collectionCode="CKC" country="Thailand" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D270FDD95791831FD10F8AFE54DF6FC" box="[755,777,1910,1936]" count="1" pageId="15" pageNumber="16" type="female">♀</specimenCount>
(
<collectionCode id="ED305C9195791831FCFAF8AFE518F6FC" box="[793,860,1910,1936]" pageId="15" pageNumber="16">CKC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B97DF9579182EFEF9F84EE35DF250" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8B9EC4549579182EFEF9F84EE35DF250" blockId="15.[188,1400,1774,2037]" lastBlockId="16.[188,1400,195,997]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="B955184695791831FEF9F84EE7D7F6DD" bold="true" box="[282,403,1943,1969]" pageId="15" pageNumber="16">Diagnosis</emphasis>
. The forewing length is 14.5–15.5 mm in males and 18.5–19.0 mm in females. 
<taxonomicName id="4C21BFD795791831FF5FF860E7B0F6BF" authorityName="Volynkin &amp; Černý" authorityYear="2025" box="[188,500,1977,2003]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cardamoma" status="sp. nov.">
<emphasis id="B955184695791831FF5FF860E7B0F6BF" box="[188,500,1977,2003]" italics="true" pageId="15" pageNumber="16">Hollowayosia cardamoma</emphasis>
</taxonomicName>
<emphasis id="B955184695791831FE1FF860E47BF6BF" bold="true" box="[508,575,1977,2003]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="A266A53D95791831FE1FF860E47BF6BF" box="[508,575,1977,2003]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131AD8D195791831FDB3F860E49DF6BF" box="[592,729,1977,2003]" captionStart="Figures 57–68" captionStartId="25.[188,274,1907,1931]" captionTargetBox="[192,1397,188,1856]" captionTargetId="figure-68@25.[188,1399,188,1898]" captionTargetPageId="25" captionText="Figures 57–68. Adults of Cernyia (57, 58), Hollowayosia gen. n. (59–62), Churingosia (63–66) and Manulea (67, 68) spp. Depositories of the specimens: 57, 67 and 68 in MfN; 58, 61 and 66 in MWM/ZSM (61: ex CKC); 59, 60 and 62 in CKC; 63–65 in NHMUK." figureDoi="http://doi.org/10.5281/zenodo.16959142" httpUri="https://zenodo.org/record/16959142/files/figure.png" pageId="15" pageNumber="16">Figs 61, 62</figureCitation>
) differs externally from 
<taxonomicName id="4C21BFD795791831FC1EF863E2CEF6BF" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[1021,1162,1977,2003]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B955184695791831FC1EF863E2CEF6BF" box="[1021,1162,1977,2003]" italics="true" pageId="15" pageNumber="16">H. longpala</emphasis>
</taxonomicName>
(
<figureCitation id="131AD8D195791831FB78F860E360F6BF" box="[1179,1316,1977,2004]" captionStart="Figures 57–68" captionStartId="25.[188,274,1907,1931]" captionTargetBox="[192,1397,188,1856]" captionTargetId="figure-68@25.[188,1399,188,1898]" captionTargetPageId="25" captionText="Figures 57–68. Adults of Cernyia (57, 58), Hollowayosia gen. n. (59–62), Churingosia (63–66) and Manulea (67, 68) spp. Depositories of the specimens: 57, 67 and 68 in MfN; 58, 61 and 66 in MWM/ZSM (61: ex CKC); 59, 60 and 62 in CKC; 63–65 in NHMUK." figureDoi="http://doi.org/10.5281/zenodo.16959142" httpUri="https://zenodo.org/record/16959142/files/figure.png" pageId="15" pageNumber="16">Figs 59, 60</figureCitation>
) in the longer forewing. Additionally, the male hindwing of the new species is paler than in the similar congener. The male genital capsule of 
<taxonomicName id="4C21BFD79566182EFD62FF1DE575F1B1" authorityName="Volynkin &amp; Černý" authorityYear="2025" box="[641,817,195,221]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cardamoma" status="sp. n.">
<emphasis id="B95518469566182EFD62FF1DE575F1B1" box="[641,817,195,221]" italics="true" pageId="16" pageNumber="17">H. cardamoma</emphasis>
</taxonomicName>
<emphasis id="B95518469566182EFCDBFF1DE53EF1B2" bold="true" box="[824,890,196,222]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A266A53D9566182EFCDBFF1DE53EF1B2" box="[824,890,196,222]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131AD8D19566182EFC6AFF1AE5A9F1B2" box="[905,1005,195,222]" captionStart="Figures 116–119" captionStartId="38.[188,274,1947,1971]" captionTargetBox="[218,1334,192,1913]" captionTargetId="figure-59@38.[202,1385,188,1928]" captionTargetPageId="38" captionText="Figures 116–119. Male genitalia of Hollowayosia gen. n. (116, 117), Churingosia nigripes (118) and Manulea palliatella (119). Depositories of the specimens dissected: 116 in CKC; 117 in MWM/ZSM; 118 in OUMNH; 119 in MfN." figureDoi="http://doi.org/10.5281/zenodo.16959188" httpUri="https://zenodo.org/record/16959188/files/figure.png" pageId="16" pageNumber="17">Fig. 117</figureCitation>
) can be easily distinguished from 
<taxonomicName id="4C21BFD79566182EFF5FFF3FE70AF193" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[188,334,229,255]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFF5FFF3FE70AF193" box="[188,334,229,255]" italics="true" pageId="16" pageNumber="17">H. longpala</emphasis>
</taxonomicName>
(
<figureCitation id="131AD8D19566182EFE87FF3CE789F193" box="[356,461,229,255]" captionStart="Figures 116–119" captionStartId="38.[188,274,1947,1971]" captionTargetBox="[218,1334,192,1913]" captionTargetId="figure-59@38.[202,1385,188,1928]" captionTargetPageId="38" captionText="Figures 116–119. Male genitalia of Hollowayosia gen. n. (116, 117), Churingosia nigripes (118) and Manulea palliatella (119). Depositories of the specimens dissected: 116 in CKC; 117 in MWM/ZSM; 118 in OUMNH; 119 in MfN." figureDoi="http://doi.org/10.5281/zenodo.16959188" httpUri="https://zenodo.org/record/16959188/files/figure.png" pageId="16" pageNumber="17">Fig. 116</figureCitation>
) by the more asymmetrical valvae (the right one is markedly broader), the narrower and apically pointed dorsal-distal section of the valva (it is apically tapered in the congener), and the longer and straighter distal saccular process with a more elongate tip. Additionally, the uncus of the new species is slightly shorter and narrower, and the vinculum is somewhat shorter than the corresponding structures in 
<taxonomicName id="4C21BFD79566182EFDEFFEB4E4D8F0EA" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[524,668,364,390]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFDEFFEB4E4D8F0EA" box="[524,668,364,390]" italics="true" pageId="16" pageNumber="17">H. longpala</emphasis>
</taxonomicName>
. The phallus of 
<taxonomicName id="4C21BFD79566182EFC85FEB4E25DF0EA" authorityName="Volynkin &amp; Černý" authorityYear="2025" box="[870,1049,364,390]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cardamoma" status="sp. n.">
<emphasis id="B95518469566182EFC85FEB4E25DF0EA" box="[870,1049,364,390]" italics="true" pageId="16" pageNumber="17">H. cardamoma</emphasis>
</taxonomicName>
<emphasis id="B95518469566182EFBC7FEB5E22DF0EA" bold="true" box="[1060,1129,364,390]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A266A53D9566182EFBC7FEB5E22DF0EA" box="[1060,1129,364,390]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is shorter and distally broader than in 
<taxonomicName id="4C21BFD79566182EFE96FE57E445F0C4" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[373,513,398,424]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFE96FE57E445F0C4" box="[373,513,398,424]" italics="true" pageId="16" pageNumber="17">H. longpala</emphasis>
</taxonomicName>
, medially downcurved (it is straight in 
<taxonomicName id="4C21BFD79566182EFC32FE57E219F0C4" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[977,1117,398,424]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFC32FE57E219F0C4" box="[977,1117,398,424]" italics="true" pageId="16" pageNumber="17">H. longpala</emphasis>
</taxonomicName>
), and bears a very short and broad, triangular carinal process, which is narrow and claw-shaped in the similar congener. The vesica of 
<taxonomicName id="4C21BFD79566182EFECEFE0BE798F087" authorityName="Volynkin &amp; Černý" authorityYear="2025" box="[301,476,465,491]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cardamoma" status="sp. n.">
<emphasis id="B95518469566182EFECEFE0BE798F087" box="[301,476,465,491]" italics="true" pageId="16" pageNumber="17">H. cardamoma</emphasis>
</taxonomicName>
<emphasis id="B95518469566182EFE07FE08E462F087" bold="true" box="[484,550,465,491]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A266A53D9566182EFE07FE08E462F087" box="[484,550,465,491]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C21BFD79566182EFD22FE0BE509F087" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[705,845,465,491]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFD22FE0BE509F087" box="[705,845,465,491]" italics="true" pageId="16" pageNumber="17">H. longpala</emphasis>
</taxonomicName>
in the broader main chamber and the markedly longer, proximally broader and granulose dorsal diverticulum, which is membranous and bearing a cluster of robust spines apically in 
<taxonomicName id="4C21BFD79566182EFDB7FDCCE4A5F343" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[596,737,533,559]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFDB7FDCCE4A5F343" box="[596,737,533,559]" italics="true" pageId="16" pageNumber="17">H. longpala</emphasis>
</taxonomicName>
. Additionally, the distal diverticulum of the new species is longer, has a broader area of graniculi, and is directed ventrally (whereas it is medially membranous and distally directed in 
<taxonomicName id="4C21BFD79566182EFE30FD80E41BF31E" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[467,607,600,626]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFE30FD80E41BF31E" box="[467,607,600,626]" italics="true" pageId="16" pageNumber="17">H. longpala</emphasis>
</taxonomicName>
), and the lateral diverticula are larger than in the similar congener. In the female genitalia, 
<taxonomicName id="4C21BFD79566182EFE32FDA3E4C5F3F8" authorityName="Volynkin &amp; Černý" authorityYear="2025" box="[465,641,634,660]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cardamoma" status="sp. n.">
<emphasis id="B95518469566182EFE32FDA3E4C5F3F8" box="[465,641,634,660]" italics="true" pageId="16" pageNumber="17">H. cardamoma</emphasis>
</taxonomicName>
<emphasis id="B95518469566182EFD6BFDA3E48EF3F8" bold="true" box="[648,714,634,660]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A266A53D9566182EFD6BFDA3E48EF3F8" box="[648,714,634,660]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131AD8D19566182EFD39FDA3E579F3F8" box="[730,829,634,660]" captionStart="Figures 147–152" captionStartId="44.[188,274,1944,1968]" captionTargetBox="[190,1369,207,1915]" captionTargetId="figure-65@44.[188,1369,188,1925]" captionTargetPageId="44" captionText="Figures 147–152. Female genitalia of Hollowayosia gen. n. (147, 148), Manulea palliatella (149) and Churingosia spp. (150–152). Depositories of the specimens dissected: 147 and 148 in CKC; 149 in MfN; 150 and 152 in NHMUK; 151 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.16959210" httpUri="https://zenodo.org/record/16959210/files/figure.png" pageId="16" pageNumber="17">Fig. 148</figureCitation>
) is distinguished from 
<taxonomicName id="4C21BFD79566182EFBAFFDA3E29CF3F8" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[1100,1240,634,660]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFBAFFDA3E29CF3F8" box="[1100,1240,634,660]" italics="true" pageId="16" pageNumber="17">H. longpala</emphasis>
</taxonomicName>
(
<figureCitation id="131AD8D19566182EFB04FDA3E30EF3F8" box="[1255,1354,634,660]" captionStart="Figures 147–152" captionStartId="44.[188,274,1944,1968]" captionTargetBox="[190,1369,207,1915]" captionTargetId="figure-65@44.[188,1369,188,1925]" captionTargetPageId="44" captionText="Figures 147–152. Female genitalia of Hollowayosia gen. n. (147, 148), Manulea palliatella (149) and Churingosia spp. (150–152). Depositories of the specimens dissected: 147 and 148 in CKC; 149 in MfN; 150 and 152 in NHMUK; 151 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.16959210" httpUri="https://zenodo.org/record/16959210/files/figure.png" pageId="16" pageNumber="17">Fig. 147</figureCitation>
) by the posteriorly broader and more asymmetrical antrum having two broad and irregular sclerotised plates (vs. one narrow and symmetrical medial plate in 
<taxonomicName id="4C21BFD79566182EFD12FD67E538F3BB" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[753,892,701,727]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFD12FD67E538F3BB" box="[753,892,701,727]" italics="true" pageId="16" pageNumber="17">H. longpala</emphasis>
</taxonomicName>
), the shorter and markedly narrower ductus bursae, the posteriorly narrower and more weakly gelatinous posterior section of the corpus bursae, the anterior section of the corpus bursae bearing two rounded and equally small signa (vs. a small rounded and a larger elliptical ones in 
<taxonomicName id="4C21BFD79566182EFDF4FCFAE4E7F250" authorityName="Volynkin &amp; Černý" authorityYear="2025" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[535,675,802,828]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFDF4FCFAE4E7F250" box="[535,675,802,828]" italics="true" pageId="16" pageNumber="17">H. longpala</emphasis>
</taxonomicName>
), and the larger and basally broader appendix bursae.
</paragraph>
</subSubSection>
<subSubSection id="C33B97DF9566182EFEF9FC9DE252F2CD" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8B9EC4549566182EFEF9FC9DE252F2CD" blockId="16.[188,1400,195,997]" pageId="16" pageNumber="17">
<emphasis id="B95518469566182EFEF9FC9DE7F0F232" bold="true" box="[282,436,836,862]" pageId="16" pageNumber="17">Distribution</emphasis>
. The new species is currently known from Southeast 
<collectingCountry id="F33684C49566182EFB85FC9DE28AF232" box="[1126,1230,836,862]" name="Thailand" pageId="16" pageNumber="17">Thailand</collectingCountry>
(
<collectingRegion id="49E50AB69566182EFB07FC9DE76EF2EC" country="Thailand" name="Chanthaburi" pageId="16" pageNumber="17">Chanthaburi Province</collectingRegion>
). The record of ‘ 
<taxonomicName id="4C21BFD79566182EFDE6FCBFE586F2EC" authority="(Holloway, 2001)" baseAuthorityName="Holloway" baseAuthorityYear="2001" box="[517,962,870,896]" class="Insecta" family="Erebidae" genus="Cernyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longpala">
<emphasis id="B95518469566182EFDE6FCBFE49DF2EC" box="[517,729,870,896]" italics="true" pageId="16" pageNumber="17">Cernyia longpala</emphasis>
(
<bibRefCitation id="EFB0B9A59566182EFD0CFCBFE5FDF2EC" author="Holloway, J. D." box="[751,953,870,896]" pageId="16" pageNumber="17" pagination="279 - 486" refId="ref17427" refString="Holloway, J. D. (2001) The Moths of Borneo, part 7. Family Arctiidae, subfamily Lithosiinae. Malayan Nature Journal, 55, 279-486." type="journal article" year="2001">Holloway, 2001</bibRefCitation>
)
</taxonomicName>
’ from 
<collectingCountry id="F33684C49566182EFBF8FCBFE2D7F2EC" box="[1051,1171,870,896]" name="Cambodia" pageId="16" pageNumber="17">Cambodia</collectingCountry>
(
<bibRefCitation id="EFB0B9A59566182EFB4AFCBFE32BF2EC" author="Bae, Y. S. &amp; Bayarsaikhan, U. &amp; Kimsun, C." box="[1193,1391,870,896]" pageId="16" pageNumber="17" refId="ref16897" refString="Bae, Y. S., Bayarsaikhan, U. &amp; Kimsun, C. (2016) Biodiversity of Cambodia. Tiger Moths (Insecta, Lepidoptera). National Institute of Biological Resources, Incheon, 303 pp." type="book" year="2016">
Bae 
<emphasis id="B95518469566182EFB06FCBEE362F2EC" box="[1253,1318,870,896]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2016
</bibRefCitation>
) probably also belongs to 
<taxonomicName id="4C21BFD79566182EFE07FC51E4D7F2CD" authorityName="Volynkin &amp; Černý" authorityYear="2025" box="[484,659,903,929]" class="Insecta" family="Erebidae" genus="Hollowayosia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cardamoma" status="sp. n.">
<emphasis id="B95518469566182EFE07FC51E4D7F2CD" box="[484,659,903,929]" italics="true" pageId="16" pageNumber="17">H. cardamoma</emphasis>
</taxonomicName>
<emphasis id="B95518469566182EFD78FC51E499F2CE" bold="true" box="[667,733,904,930]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A266A53D9566182EFD78FC51E499F2CE" box="[667,733,904,930]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
but requires confirmation.
</paragraph>
</subSubSection>
<subSubSection id="C33B97DF9566182EFEF9FC70E2F6F289" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8B9EC4549566182EFEF9FC70E2F6F289" blockId="16.[188,1400,195,997]" pageId="16" pageNumber="17">
<emphasis id="B95518469566182EFEF9FC70E7E6F2AF" bold="true" box="[282,418,937,963]" pageId="16" pageNumber="17">Etymology</emphasis>
. The specific epithet is derived from the Cardamom (Khelang) Mountains, where the new species is found. The name is a noun in the nominative singular in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>