Hanken, James, Wake, David B. & Savage, Jay M., 2005, A solution to the large black salamander problem (genus Bolitoglossa) in Costa Rica and Panamá.

Hanken, James, Wake, David B. & Savage, Jay M., 2005, A solution to the large black salamander problem (genus Bolitoglossa) in Costa Rica and Panamá., Copeia 2005, pp. 227-245

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