Platyscelis (Platyscelis) brevis Baudi di Selve, 1876

Bai, Xing-Long & Ren, Guo-Dong, 2019, Revision of the genus Platyscelis Latreille, 1818 from China (Coleoptera: Tenebrionidae: Platyscelidini), Zootaxa 4609 (1), pp. 101-126 : 108-111

Platyscelis (Platyscelis) brevis Baudi di Selve, 1876

( Figs. 15–16 , 47–49 , 68 )

Platyscelis brevis Baudi di Selve, 1876: 35 ; Kaszab, 1940: 927 (synonym of Blaps rugifrons Germar, 1823 ).

Platyscelis (Platyscelis) brevis: Egorov, 1990: 891 ; 2004: 661; 2008: 296; 2009: 67; Ren & Yu, 1999: 304; Yu et al., 1999: 365

(pupae); Yu et al., 2000: 163 (larvae); Ren et al., 2000: 4 (list of larvae); Ren & Liu, 2009: 1149 (defensive gland).

Blaps rugifrons Germar, 1823: 139 (synonymized by Egorov, 1990: 891).

Platyscelis rugifrons: Fischer von Waldheim, 1824: 194 ; Solier, 1848: 210; Seidlitz, 1893: 345.

Platyscelis (Platyscelis) rugifrons: Kaszab, 1940: 927 ; Egorov, 1989: 346, 350.

Platyscelis ballioni Reitter, 1889: 698 , syn. n.

Platyscelis ballioni: Seidlitz, 1893: 345 .

Platyscelis (Platyscelis) ballioni: Kaszab, 1940: 929 ; Egorov, 1989: 350; 2004: 661; 2008: 296; 2009: 67.

Type material (studied). Paralectotype of Platyscelis ballioni : 1♀ ( HNHM), Shan-si, 1884, G. Patanin // Platyscelis ballioni m. 1889 // Typus, Platyscelis ballioni Reitter, Coll. Reitter // Platyscelis ballioni Reitt. , det. L. Egorov 1989 ( Figs. 47 , 68 ) .

Other material. CHINA: 1♀ ( IZCAS), Manzha , 24.XI.75; Hebei : 7♂♂, 2♀♀ ( MHBU), Guyuan , 24.VII.1998, Ming Bai leg. ; 1♂ ( MHBU), Tunken Forest Farm, Kangbao , 1.VIII.1998, Ming Bai leg. ; 3♀♀ ( MHBU), Tunken, Kangbao , 41°57′38.8″ N, 114°44′23.3″ E, 1496 m, 24.VII.2012, Yi-Ping Niu & Xiao-Jie Sun leg. ; 1♀ ( MHBU), Fengning , 24.VII.2002, Gui-Ming Tang leg. ; 2♂♂ ( MHBU), Suangou, Shangyi , 29.VI.2003, Hui Zhen leg. ; 18♂♂, 10♀♀ ( MHBU), Yudaokou, Chengde , 12–18.VI.2004, Chao Gao leg. ; 1♂ ( MHBU), Xiaoquangou, Chazi, Zhongmiaochang , Mulanweichang , Weichang , 42°05′34.9″ N, 117°25′05.5″ E, 1288 m, 27.V.2015, Li Ma leg. ; 1♀ ( MHBU), Chazi, Zhongmiaochang, Mulanweichang , Weichang , 41°43′02.6″ N, 118°02′01.4″ E, 1078 m, 27.VI.2015, Di Li leg. ; 1♀ ( MHBU), Liangtou, Wudaogou, Mulanweichang , Weichang , 41°55′16.0″ N, 116°50′13.9″ E, 1720 m, 30.VI.2015, Di Li leg. ; 2♀♀ ( MHBU), Yanboge, Mulanweichang, Weichang , 42°10′19.6″ N, 117°17′47.8″ E, 1380 m, 14.VII.2016, Jing-Jing Ma & Li Ma leg. ; 1♂ ( MHBU), Bashihao, Dahuanqi, Saihanba , Weichang , 42°09.146′ N, 117°25.351′ E, 1189 m, 1.VI.2015, Jun-Sheng Shan & Cheng Fang et al. leg .; 1♂ ( MHBU), Baishui, Yinhe, Saihanba , Weichang , 42°22.243′ N, 117°26.723′ E, 1510 m, 26.VI.2015, Jun-Sheng Shan & Cheng Fang et al. leg .; 1♂ ( MHBU), Yinhe, Saihanba, Weichang , 42°19′4.2″ N, 117°31′37.3″ E, 1672 m, 10.VIII.2016, Jian-Bo Zhou leg. ; 11♂♂, 6♀♀ ( MHBU), Yangjiaping, Zhuolu , 25–29.VI.2009, Xin-Pu Wang & Zhen-Hua Gao et al. leg.; Heilongjiang : 1♂ ( IZCAS), Liangshuigou, Dailing , [Yichun], 26.V.1957; Inner Mongolia : 1♂, 1♀ ( MHBU), Zhengxiangbai Banner , 14.VIII.1978, He-Ming Chen leg. ; 1♂, 1♀ ( MHBU), Zhengxiangbai Banner , 1450 m, 20.IV.1991, Yuan-Chao Guo leg. ; 1♂, 1♀ ( MHBU), Zhengxiangbai Banner , 14.VIII.1991, He-Ming Chen leg. ; 1♂ ( MHBU), Zhengxiangbai Banner , 1.V.2000, Yuan-Chao Guo leg. ; 2♂♂, 3♀♀ ( MHBU), Zhengxiangbai Banner , 25.V.2002, Yuan-Chao Guo leg. ; 1♀ ( MHBU), Zhengxiangbai Banner , 19.VI.2004, Yuan-Chao Guo leg. ; 1♀ ( MHBU), Xingyao, Zhengxiangbai Banner , 1250 m, 21.V.1984, Yuan-Chao Guo leg. ; 2♀♀ ( MHBU), Ming’antu, Zhengxiangbai Banner, Xilingol League , 1345 m, 30.V.2013, Yuan-Chao Guo leg. ; 3♂♂, 1♀ ( MHBU), Ming’antu, Zhengxiangbai Banner, Xilingol League , 1345 m, 9.VI.2013, Yuan-Chao Guo leg. ; 1♂, 1♀ ( MHBU), Ming’antu, Zhengxiangbai Banner, Xilingol League , 1385 m, 1.VI.2013, Yuan-Chao Guo leg. ; 11♂♂, 18♀♀ ( MHBU), Ming’antu, Zhengxiangbai Banner, Xilingol League , 1360 m, 10.VI.2013, Yuan-Chao Guo leg. ; 1♀ ( MHBU), Ming’antu, Zhengxiangbai Banner, Xilingol League , 1360 m, 19.VI.2013, Yuan-Chao Guo leg. ; 4♂♂ ( KIZ), Manzhouli , 6.VII.1981, Jiang Xiong leg. ; 12♂♂, 4♀♀ ( MHBU), Xilingol League , 4.IV.1987, Yuan-Chao Guo leg. ; 1♂ ( MHBU), Alxa Left Banner , 15.VIII.1984, Qiang Liu leg. ; 1♂ ( MHBU), Yabrai Forest Farm, Alxa Right Banner , 15.VIII.1984, Qiang Liu leg. ; 2♂♂ ( MHBU), Erdaogou, Jiufeng Shan, Baotou , 9.V.1985, Chang-Qing Mao leg. ; 1♀ ( MHBU), Erdaogou, Jiufeng Shan , 9.V.1985, You-Zhi Yu leg. ; 9♂♂, 1♀ ( MHBU), Bayan Obo, Baotou , 3.VIII.1991, Guo-Dong Ren leg. ; 1♂ ( MHBU), West Ujimqin Banner , 14.VI.1994, Guo-Dong Ren leg. ; 13♂♂, 18♀♀ ( MHBU), East Ujimqin Banner , 15.VI.1994, Guo-Dong Ren leg. ; 1♂ ( MHBU), East Ujimqin Banner , 5.VII.1994, Guo-Dong Ren leg. ; 1♀ ( MHBU), Balamawula, East Ujimqin Banner, Xilingol League , 46°34.053′ N, 119°14.962′ E, 1092 m, 15.VII.2014, Hao-Yu Liu, Zhao Pan, Xiao-Jie Sun & Yan Yan leg. ; 7♂♂, 4♀♀ ( MHBU), Horqin , 14.VI.1994, Guo-Dong Ren leg. ; 1♀ ( MHBU), Bayan Huxu, Horqin , 14.VIII.1997, Guo-Dong Ren leg. ; 2♂♂, 2♀♀ ( MHBU), Hexigten Banner , 18.VI.1994, Guo-Dong Ren leg. ; 1♂ ( MHBU), Hexigten Banner , Qiang Liu leg. ; 1♀ ( MHBU), Ongniud Banner , 27.VI.1994, Guo-Dong Ren leg. ; 1♂ ( MHBU), Qigan, Ongniud Banner, Chifeng , 42°55.727′ N, 119°18.736′ E, 619 m, 17.VII.2014, Hao-Yu Liu, Zhao Pan, Xiao-Jie Sun & Yan Yan leg. ; 5♂♂, 9♀♀ ( MHBU), Yezhulin, Bairin Left Banner, Chifeng , 43°45.409′ N, 119°20.387′ E, 619 m, 27.VII.2014, Hao-Yu Liu, Zhao Pan, Xiao-Jie Sun & Yan Yan leg. ; 3♂♂, 2♀♀ ( MHBU), Xin Barag Right Banner, 23.VII.2004, Guo-Dong Ren & Wen-Jun Hou leg. ; 5♂♂, 6♀♀ ( MHBU), Xin Barag Left Banner , 25.VIII.2008, Fu-Ming Shi leg. ; 1♂, 2♀♀ ( MHBU), Xin Barag Left Banner , 26.VIII.2008, Fu-Ming Shi leg. ; 1♂ ( MHBU), ca. 4 km NE of Xin Barag Left Banner, Hulun Buir , 48.18269° N, 118.29554° E, 717 m, 17.VII.2014, Hao-Yu Liu, Zhao Pan, Xiao-Jie Sun & Yan Yan leg. ; 4♂♂, 4♀♀ ( MHBU), 6 km W of Ergun , 26.VII.2004, Guo-Dong Ren & Wen-Jun Hou leg. ; 2♂♂, 1♀ ( MHBU), Eerdunwul agacha, Chen Barag Banner, Hulun Buir , 49°24.939′ N, 118°43.010′ E, 588 m, 19.VII.2014, Hao-Yu Liu, Zhao Pan, Xiao-Jie Sun & Yan Yan leg. ; 1♀ ( MHBU), Hadat, Chen Barag Banner, Hulun Buir , 49°40.515′ N, 119°48.021′ E, 674 m, 19.VII.2014, Hao-Yu Liu, Zhao Pan, Xiao-Jie Sun & Yan Yan leg. ; 4♂♂, 2♀♀ ( MHBU), Chaoyang, Huade , 30.IV.2008, Guo-Dong Ren, Ben-Yong Mao & Wei Li leg. ; 2♂♂, 2♀♀ ( MHBU), Changhanliang, Ulanqab , 40°46′58.9″ N, 112°49′16.7″ E, 1663 m, 23.VII.2012, Yi-Bin Ba & Yi-Ping Niu leg. ; 1♂, 1♀ ( MHBU), Chahanying, Zhuozi , 27.VI.2018, Guo-Dong Ren, Yi-Bin Ba & Ming-Min Ma leg. , 1♂, 1♀ ( MHBU), Xilinhot , 18.VII.2009, Qing-Wei Huo leg. ; 5♂♂, 2♀♀ ( MHBU), Bayan Qagan, Chahar Right Back Banner , 41°26′41.0″ N, 113°17′29.4″ E, 1480 m, 23.VII.2012, Yi-Bin Ba & Yi-Ping Niu leg. ; Jilin: 1♂ ( IZCAS), Pitai , 28.V.1957; Liaoning : 1♀ (IZCAS), Zhanggutai , [Zhangwu], 19.V.1956; Shandong : 4♂♂, 2♀♀ ( IZCAS), North-Eastern [in Chinese] // Ta-Yngtse, Linsisien [Linzi, Shangdong], leg E. Bourgault, VII.1939 ; 1♂, 2♀♀ ( IZCAS), North-Eastern [in Chinese] // Ta-Yngtse, Linsisien , leg E. Bourgault, VII.40 ; 1♂ ( IZCAS), North-Eastern [in Chinese] // Ta-Yngtse, Linsisien , leg E. Bourgault, VI.40; Shanxi : 1♀ ( HNHM), Shoachow, China, Shansi // Platyscelis ballioni Reitter , det. dr. Kaszab // Platyscelis ballioni Reitt. , det. L. Egorov 1989 . MONGOLIA: 1♀ ( HNHM), Mongolia, Chara-gol // Exp. Zichy, leg. Csiki // coll. Dr. J. Fodor // det. L. Egorov, 1990; 2♂♂, 2♀♀ ( MHBU), Khan Khen National Park , 11.VIII.2001, Gui-Ming Tang leg. ; 1♀ ( MHBU), Bürd , Ovorhangay, 47°06′44.8″ N, 103°36′58.3″ E, 1467 m, 22.VII.2015, Xiao-Jie Sun leg. RUSSIA: 1♂ ( HNHM), Tschita , Coll. Lichtn. // det. L. Egorov, 1990; 2♀♀ ( IZCAS), North-Eastern [in Chinese] // Tschita été 192, leg. V.J.Tolmachov, Musée Heude.

Distribution. China: Hebei ( Kaszab, 1940; Liu et al., 2015), Heilongjiang (new record), Inner Mongolia ( Kaszab, 1940; Ren & Yu, 1999; Egorov, 2008), Jilin (new record), Liaoning (new record), Shandong (new record), Shanxi ( Reitter, 1889; Kaszab, 1940; Egorov, 1989, 2008); Mongolia ( Kaszab, 1940; Ren & Yu, 1999; Egorov, 2008); Russia ( Kaszab, 1940; Egorov, 2008; Dorzhieva, 2010).

Remarks. Platyscelis ballioni ( Fig. 47 ) was published by Reitter (1889) with a single female specimen from Shanxi, China. Kaszab (1940) examined and redescribed the specimen in his revision of the tribe Platyscelidini worldwide, and diagnosed it from Blaps rugifrons Germar with the characters: elytra quite flat and parallel (arched and short oval in B. rugifrons ); apex of protibiae strongly foliate extended (not strongly in B. rugifrons ). P. brevis ( Figs. 48–49 ) distributed in Mongolian Plateau, 291 specimens were examined in this study, body short in north and longer in south generally. Therefore, P. ballioni was a female of P. brevis in southern Mongolian Plateau.

Figures

FIGURES 5–20. Parameres of male aedeagus in dorsal and lateral view. 5–6. Platyscelis acutipenis; 7–8. P. amdoensis; 9–10. P. angusticollis angusticollis; 11–12. P. angusticollis platytarsis; 13–14. P. bogatshevi; 15–16. P. brevis; 17–18. P. confusa;

FIGURES 45–50. Habitus. 45. Platyscelis angusticollis platytarsis (male); 46. P. bogatshevi (male); 47–49. P. brevis (47. female, paralectotype of P. ballioni; 48. female; 49. male); 50. P. confusa (male, lectotype).

FIGURES 63–72. Habitus and type labels. 63–65. Habitus: 63. Platyscelis suiyuana (male); 64. P. ovata (male); 65. P. striata (male). 66–72. Type labels: 66. P. angusticollis angusticollis; 67. P. angusticollis platytarsis; 68. P. ballioni; 69. P. hauseri (= P. confusa); 70. P. gebieni; 71. P. licenti; 72. P. suiyuana.