Pheidole kugleri, Wilson, E. O., 2003

Wilson, E. O., 2003, Pheidole in the New World. A dominant, hyperdiverse ant genus., Cambridge, MA: Harvard University Press : 309

Pheidole kugleri new species

Types Mus. Comp. Zool. Harvard.

etymology Named after Charles Kugler, in recognition of his research in myrmecology and his extraordinary collections of Pheidole made in Colombia and other parts of South America.

diagnosis A member of the " jujuyensis complex" of the larger fallen group, comprising araneoides , cuevasi , durionei , jujuyensis , kugleri , leonina , leptina , lucretii , lupus , paraensis , punctithorax , tijucana , wallacei , and wolfringi . P. kugleri is easily distinguished from the other species by the following traits.

Major: antennal scapes exceed occipital corners by twice their maximum width; carinulae originating on frontal lobes and along frontal carinae spread laterally out to create a fan-like pattern, with those coming from the more anterior parts of the frontal carinae turning almost horizontal to travel toward the sides of the head; rugoreticula on each side of the head extend from the eye to the antennal fossa; occiput narrowed, the nadir of its cleft deep and rounded; promesonotum and all surfaces of the propodeum carinulate. The holotype is more heavily sculptured than the average. Minor: unknown.

Measurements (mm) Holotype major: HW 1.34, HL 1.50, SL 1.34, EL 0.24, PW 0.68. color Major: body dark brown, appendages medium brown.

Range Known only from the type locality.

biology The type series was collected from primary forest on a steep hillside above farmland.

Figure Holotype, major. PERU: 3 km north of Oxacampa, Pasco, 2000 m (Charles Kugler and R. R. Lambert). No minors available. Scale bar = 1 mm.