Phanoperla batac, Sivec & Stark, 2011

Sivec, Ignac & Stark, Bill P., 2011, New Species Of Neoperla Needham And Phanoperla Banks (Plecoptera: Perlidae) From The Philippine Islands, Illiesia 7 (24), pp. 264-279 : 273-274

Phanoperla batac sp. n.

( Figs. 35-44 )

Material examined. Holotype ♂ and 1♀ paratypes, PHILIPPINE ISLANDS, Palawan, San Rafael, Ulanguan, Batac Village , 10 March 1996, (Holotype PZ), 1♀ paratype ( PMSL). Additional paratypes: Palawan, San Rafael , Ulanguan , end of February 1996, 5♂ ( PZ), 1♂ ( PMSL). Palawan, Salakot Falls Road , 19 March 1996, 4♂, 1♀ ( PZ). Palawan, Mt. Salakot , Heli-Pad , 720 m, 19 March 1996, 1♂ ( PZ). Specimens deposited in the PZ collection, or the PMSL as indicated.

Adult habitus. General color pale yellow or white. Head uniformly pale, ocelli small, about two diameters apart; callosities and M-line indistinct, antennae and palpi pale but 1 st antennal segment brownish. Pronotum pale but slightly darker along margins and median suture ( Fig. 39 ). Legs and wings pale.

Male. Forewing length 9.5 mm. Tergum 9 bearing a large mesal patch of sensilla narrowly divided on midline ( Fig. 35 ). Hemiterga with a slightly raised, elongate swelling overlapping base of hemitergal lobes; hemitergal lobes excavated along inner margins near apex, relatively straight and directed mesad. Abdominal sterna 4-5 with ventral hair brushes. Penis tube poorly sclerotized but bearing an irregular dorsal sclerite and a thumb shaped, apically directed spiny lobe; penis sac armed only near apex on lateral and dorsal surface but with a narrow midventral band of spines ( Figs. 36-38 ); subapex bearing an almost complete ring of ca. 14 larger black spines; apex of sac abruptly narrowed beyond spine ring and armed with a dense coating of small black spines; dorsum of sac bearing a small spiny lobe near apex.

Female. Forewing length 10-12 mm. Subgenital plate poorly developed, not projecting beyond posterior margin of sternum 8.

Egg. Outline tear drop shaped with long, slender collar, rugose along surface and with a small flanged rim ( Fig. 40-44 ); chorionic surface with a median zone of fine punctations and a small zone of coarser punctations on lid.

Etymology. The species name, used as a noun in apposition, honors the Batac people of Palawan.

Diagnosis. The penis tube and sac of this species is generally similar to that of P. sertispina Jewett, 1975 , an Indian member of the P. anomala ( Banks, 1939) species group. This species shares a relatively complete apical ring of large black penial spines, other sac armature limited primarily to the apical half, and a tear drop shaped egg with long slender collar with members of the P. anomala complex, and should probably be assigned to that group. The new species differs from these species in having an almost cylindrical and relatively straight penial tube and sac, and in having the dorsal thumb shaped lobe somewhat larger and covered with small spines.

Figures

Figs. 35-40. Phanoperla batac. 35. Male terminalia. 36-38. Penis tube and everted sac. 36. Dorsal aspect. 37. Ventral aspect. 38. Lateral aspect. 39. Head and pronotum. 40. Egg.

Figs. 41-44. Phanoperla batac egg. 41. Entire egg. 42. Chorionic detail. 43. Anterior end. 44. Collar end.