Hoplopholcus labyrinthi ( Kulczyński, 1903 )

Huber, Bernhard A., 2020, Revision of the spider genus Hoplopholcus Kulczyński (Araneae, Pholcidae), Zootaxa 4726 (1), pp. 1-94 : 9

Hoplopholcus labyrinthi ( Kulczyński, 1903)

Figs 6–7 , 12–46

Holocnemus labyrinthi Kulczyński, 1903: 44 , pl. 1, figs 5–7 (♂ ♀).

Artema cretica Roewer, 1928: 121 , figs 31–34 (♂ ♀). Synonymized by Senglet (1971).

Neartema cretica — Kratochvíl 1940: 6 .

Hoplopholcus labyrinthi — Senglet 1971: 348 , figs 11–16. Brignoli 1976: 559. Brignoli 1984: 289. Gasparo 2003: 59. Gasparo 2006: 59. Bosmans et al. 2013: 25 (except specimens from Milatos Cave, Katofygi Caves, Koufeospileos Cave, and from Karouba; see H. minotaurinus ). Naumova et al. 2016: 434. Eberle et al. 2018 (molecular data). Huber et al. 2018: fig. 6.

Pholcus opilionoides (Schrank) (misidentification)— Roewer 1959: 10 (most of the specimens listed from Crete; see Notes below).

Hoplopholcus minotaurinus (misidentifications)— Brignoli 1976: 561 (only specimens from “Grotta Ilithia”). Bosmans et al. 2013: 26 (only specimens from Doxa Cave, Kronos Antron Cave, and Trapezas Cave).

Figures

FIGURES 6–11. Hoplopholcus spp., live specimens. 6–7. H. labyrinthi (Kulczyński, 1903), male and female from Spilaio Melidoniou. 8–9. H. suluin sp. n., male and female from Sulu İn Mağarası. 10–11. H. minous Senglet, 1971, male from Orino Gorge and female with egg-sac from Exo Mouliana.

FIGURES 12–20. Hoplopholcus labyrinthi (Kulczyński, 1903). Left male palp in prolateral, dorsal, and retrolateral views (ar- rows in Fig. 12 point at femur-patella joints; arrow in Fig. 14 points at transversal dark line on femur); epigyna, ventral views, and cleared female genitalia, ventral and dorsal views. 12–17. Male and female from Spilaio Melidoniou (ZFMK Ar 20875). 18–20. Female from Spilaio Vreikou (ZFMK Ar 20884).

FIGURES 21–30. Hoplopholcus labyrinthi (Kulczyński, 1903) from Spilaio Melidoniou (ZFMK Ar 20875). 21–22. Left pro- cursus, prolateral and retrolateral views (arrow points at distinctive membranous process). 23–24. Male chelicerae, frontal and lateral views. 25–28. Left genital bulb, dorsal, prolateral, ventral, and retrolateral views (arrow points at distinctive prolateral process). 29–30. Cleared female genitalia, ventral and dorsal views. dmp, dorsal membranous process; es, embolar sclerite; mp, median pouch of uterus externus; lp, lateral pouch of uterus externus; tp, transparent process; vk, ventral ‘knee’; vs, ventral sclerite. Scale lines: 0.5 mm.

FIGURES 31–36. Hoplopholcus labyrinthi (Kulczyński, 1903) (ZFMK Ar 20875). 31–32. Male and female carapace and ocu- lar area, frontal (slightly dorsal) views. 33–34. Stridulatory ridges on male and female chelicerae. 35. Tip of left male cheliceral apophysis with cone-shaped hairs. 36. Male gonopore with epiandrous spigots. Scale lines: 10 µm (35), 20 µm (33), 100 µm (31–32, 34, 36).

FIGURES 37–42. Hoplopholcus labyrinthi (Kulczyński, 1903) (ZFMK Ar 20875). 37–38. Male and female palpal tarsal or- gans. 39. Tip of right procursus, retrolateral view, showing ventral spine (vs), transparent process (tp), and distinctive membranous process (arrow). 40. Hair-like processes at tip of procursus. 41. Tip of embolar sclerite with hair-like processes. 42. Base of trichobothrium on female palpal tibia. Scale lines: 10 µm (37–38, 40–42), 100 µm (39).

FIGURES 43–46. Hoplopholcus labyrinthi (Kulczyński, 1903) (ZFMK Ar 20875). 43–44. Male and female ALS. 45. Spines on male femur 1. 46. Curved hairs on male metatarsus 2. Scale lines: 10 µm (43–44), 20 µm (45), 100 µm (46).