Dolichognatha incanescens (Simon)

Smith, Helen M., 2008, Synonymy of Homalopoltys (Araneae: Araneidae) with the genus Dolichognatha (Araneae: Tetragnathidae) and descriptions of two new species, Zootaxa 1775, pp. 1-24 : 15-18

publication ID

https://doi.org/ 10.5281/zenodo.182204

DOI

https://doi.org/10.5281/zenodo.6233876

persistent identifier

https://treatment.plazi.org/id/B80987DF-4F7B-FFFE-0399-FD72FB4C05B5

treatment provided by

Plazi

scientific name

Dolichognatha incanescens (Simon)
status

 

Dolichognatha incanescens (Simon) View in CoL

Figs 1–16 View FIGURES 1 – 7 View FIGURES 8 – 12 View FIGURES 13 – 16 , 38–53 View FIGURES 38 – 45 View FIGURES 46 – 53

Homalopoltys incanescens Simon, 1895: 893 . Holotype Ψ, Sri Lanka, Galle. In MNHNP, No.16311, examined. NEW COMBINATION.

Other material examined. SRI LANKA: ɗ, RMNH.ARA.11302 (ex coll. CLD), Ratnapura, 21–22.viii.1981, forest & lake below tennis club. AUSTRALIA: Queensland: 2 Ψ, QM S73921, Bellenden Ker Range, Cableway base stn, 17–24.x.1981, Earthwatch/Qld Museum, 100 m; 2 ɗ 1 Ψ, QM S74359 View Materials , Gordon Creek, Iron Range, 24–30.vi.1976, R. Raven, V.E. Davies; 1 ɗ 1 Ψ, AM KS 100890, data as S74359 View Materials ; ɗ, QM S74358 View Materials , Spear Creek, 3–10.xi.1975, R. Raven, V.E. Davies; 2ɗ, QM S74356 View Materials –7, Wallaman Falls, via Ingham, 1.x.1980, G. Monteith, 500 m RF, pyrethrum. PAPUA NEW GUINEA: 4ɗ 20Ψ, RBIN, Baiteta Forest, 5°01'S 145°45'E, 1993–96, canopy fogging. Material of doubtful identity: INDONESIA: Kalimantan: 2 Ψ, RMNH (ex coll. CLD), Tumbang Tahai, 2°02'S 113°35'E, 3–11.ix.85, Suh. Djojosudharmo, primary moist forest; Ψ, RMNH (ex coll. CLD), Kaharian, 2°02'S 113°40'E, 2–16.ix.85, Suh. Djojosudharmo, swampy primary forest, foliage.

Comments. The female discussed here as the type of H. incanescens is believed to be that of E. Simon (1895). As discussed under D. albida , the type locality in the specimen vial does not match that given in the original publication, where it is Kandy. There are two registration numbers with the H. incanescens specimen. The number used by MNHNP on the loan form is given above. The middle digit on the other label (probably Simon’s label, which also carries the locality “Galle!”) is not clear, but looks more like a ‘2’.

Diagnosis. Male. Palpal bulbus almost twice as long as wide, embolus without spine-like apophysis ( Figs 43, 44 View FIGURES 38 – 45 ).

Female. Abdomen bilobed apically in dorsal view ( Fig. 47 View FIGURES 46 – 53 ), strongly constricted below apical tubercles; epigynum posterior margin indented medially in ventral view ( Fig. 50 View FIGURES 46 – 53 ).

Description. Male (RMNH.ARA.11302). Carapace: length 1.00 (range 0.88–1.08), width 0.82, height 0.55; strongly humped in lateral view ( Fig. 38 View FIGURES 38 – 45 ), little differentiation between caput and posterior part (some other males more similar to other species), caput sides sloping in dorsal view ( Fig. 40 View FIGURES 38 – 45 ), anterior margins of carapace squared, but with small protrusions in some New Guinea males. Chelicerae ( Figs 38, 42 View FIGURES 38 – 45 ): small, angled cheliceral ledge; two distinct promarginal cheliceral teeth, 6 (right) or 5 (left) medium and small teeth scattered intermediately and on retromargin (right side with three small teeth apparently in a transverse row across groove), other males equally variable, one appears to have 5 retromargin teeth on one side but none at all on the other. Labium: labial spur strongly produced ( Fig. 38 View FIGURES 38 – 45 ). Sternum ( Figs 38, 41 View FIGURES 38 – 45 ): slightly convex, but less so than in D. deelemanae . Eyes ( Figs 38, 40, 42 View FIGURES 38 – 45 ). AME: 0.10, PME: 0.06, ALE: 0.075, PLE: 0.065, AME–AME: 0.03, AME–ALE: 0.035, PME–PME: 0.03, PME–PLE: 0.07, ALE–PLE: 0.01; outline of tapeta clearly visible in secondary eyes. Legs ( Fig 38 View FIGURES 38 – 45 ): I: 4.75, II: 4.04, III: 2.10, IV: 2.53; anterior femurs with three dorsolateral macrosetae distally, 2 prodorsal, 1 retrodorsal. Abdomen ( Figs 38, 39 View FIGURES 38 – 45 ): length 1.45, width 0.76; apex rounded; post-genital mound not developed. Palpal organ ( Figs 13–16 View FIGURES 13 – 16 , 43–45 View FIGURES 38 – 45 ): palpal femur 0.29× carapace length; patella without macroseta, tibia c. 2× as long as wide; embolus gently tapering to blunt tip, embolic apophysis spoon-shaped distally, stem longer and more strap-like than D. mandibularis ; conductor, embolus and apophysis all more elongate than in D. mandibularis ; bulbus width: length = 1:1.8. Colour in alcohol: almost entirely pale creamy white and semi translucent in parts; PME with black around margins; faint trace of inverted arrow-head on sternum ( Fig. 41 View FIGURES 38 – 45 ); dorsal abdomen with traces of dark pigment posteriorly.

Female (holotype). Carapace: length 1.25 (range 1.00–1.25), width 1.02, height 0.63; in lateral view rather similar to male with caput poorly differentiated from posterior ( Fig. 46 View FIGURES 46 – 53 ), rather broad posteriorly in dorsal view, caput region short ( Fig. 48 View FIGURES 46 – 53 ); some other females are more similar in carapace shape to other species. Chelicerae: promargin with two definite teeth plus large intermediate where rows meet ( Figs 9, 10 View FIGURES 8 – 12 , intermediate arrowed), retromargin with one large and 4 small teeth grouped distally ( Fig. 10 View FIGURES 8 – 12 ). Labium: rounded ( Fig. 8 View FIGURES 8 – 12 ), mounded on anterior face. Maxillae with serrula ( Fig. 8 View FIGURES 8 – 12 ). Sternum: convex, but less pronounced than in D. deelemanae . Eyes ( Figs 46, 48 View FIGURES 46 – 53 ): AME: 0.11, PME: 0.09, ALE: 0.09, PLE: 0.08, AME–AME: 0.035, AME– ALE: 0.05, PME–PME: 0.03, PME–PLE: 0.06, ALE–PLE: 0.015; no reflective tapeta present in type, but visible in all secondary eyes of several recently collected females from New Guinea and Queensland; lateral eyes are close in all specimens, but eye position varies both vertically and horizontally. Legs ( Fig. 46 View FIGURES 46 – 53 ): I: 4.75, II: 4.20, III: 2.22, IV: 2.78; distal macrosetae missing, no indication of any prolateral macrosetal bases. Abdomen ( Figs 46, 47 View FIGURES 46 – 53 ): length 2.53, width 1.39; apically rather more constricted than D. raveni , and many New Guinea specimens are narrower still; post genital mound well developed as a rounded bulge, but less distinct in some other specimens. Epigyne ( Figs 50, 51 View FIGURES 46 – 53 ): posterior margin broadly indented, copulatory openings hidden in ventral view; in posterior view the internal margins of the lateral plates converge towards the copulatory openings. Internal genitalia ( Fig. 52 View FIGURES 46 – 53 ): copulatory ducts pass spermathecae medially, entering posteromedially into lower lobe of spermatheca; spermatheca head globular-ovate. Colour in alcohol: carapace creamy-white, eye region and chelicerae with olive markings, sternum with dark mark. Legs cream with olive-brown annulations, incomplete on femora. Abdomen dorsum pale except for brown patch anterior to anal tubercle and remains of paired markings towards anterior; brown lateral patches; ventrally with olive-brown over post-genital mound, outlined with white lines. The New Guinea females consistently have a slightly different carapace pattern with more brown or black on the caput ( Fig. 49 View FIGURES 46 – 53 ) and the brown bar anterior to the spinnerets on the dorsal abdomen becomes black laterally, persisting as black spots in nearly every specimen. The Queensland females are either bleached or in poor condition, but appear to more closely match the holotype colour pattern.

Specimens of doubtful identity. The female specimens examined from Kalimantan have rather shorter copulatory ducts ( Fig. 53 View FIGURES 46 – 53 ), no male is available from this area.

Distribution. Dolichognatha incanescens occurs from Sri Lanka to north-eastern Australia.

SRI

Serengetti Research Institute

RMNH

National Museum of Natural History, Naturalis

CLD

Cleveland Literary and Philosophical Society

NEW

University of Newcastle

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Tetragnathidae

Genus

Dolichognatha

Loc

Dolichognatha incanescens (Simon)

Smith, Helen M. 2008
2008
Loc

Homalopoltys incanescens

Simon 1895: 893
1895
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