Salcedia utetea, Balkenohl, 2020

Balkenohl, Michael, 2020, A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini), ZooKeys 901, pp. 1-81 : 1

publication ID	https://dx.doi.org/10.3897/zookeys.901.39432
publication LSID	lsid:zoobank.org:pub:843F6065-D00E-48E6-9A26-9CEB044BF102
persistent identifier	https://treatment.plazi.org/id/33502FE5-EE6E-4405-86AD-BC8FE93E1A6D
taxon LSID	lsid:zoobank.org:act:33502FE5-EE6E-4405-86AD-BC8FE93E1A6D
treatment provided by	ZooKeys by Pensoft
scientific name	Salcedia utetea
status	sp. nov.

Treatment

Salcedia utetea View in CoL sp. nov. Figs 20 View Figures 20–25 , 38 View Figures 36–43 , 64 View Figures 64–67 , 79 View Figures 77–83 , 84 View Figure 84

Type material.

Holotype: ♂, with labels and data: white, black printed "TANZANIA or.mer. 8°03,2'S, 38°52,7'E; 70m 15 km N Utete, pr.Pwani leg. F. Kantner 16.i.2007" (SMNS). Paratypes: 1 ♂, 1 ♀, 12 specs., same data as holotype (SMNS, CBB); 1 ♂, 3 specs., same data as holotype (CBP); 1 spec., "Tanzania prov. Pwani, 15 km E Utete, 75 m, 16.1.2007, leg. Halada / Salcedia sp. det. Schuh 2009 / Salcedia spec. P. Schüle det. 2012" (CBP); 1 ♂, with labels and data: beige, black printed and handwritten with black ink "Narobi b. Tanga D. O. Afr. V 1915 leg. Methner" / "COM. INST. ENT. COLL. NO. 13458" (MFNB); 1 ♂, same data as holotype but first label handwritten “Ksagara” (MFNB).

Diagnosis.

A medium sized species, with sub-elongate outline of the elytra with maximum width at middle and the pronotum with complete inner and outer lateral carinae. The pseudohumerus is sub-rectangular with a laterally projecting tooth. The antennomeres are elongate. Distinguished most clearly from the similar species S. nigeriensis by the lateral margin of the pronotum with ten tubercles and the complete set of carinae on the disc. In addition and in contrast to S. nigeriensis , the lateral margin of the elytron (not visible in dorsal view) is smooth.

Description.

Measurements in Table 1 View Table 1 .

Colour and surface: Piceous, shiny; top of carinae on head, pronotum and elytra as well as margins of pronotum and elytra opaque, covered with pale grey pili; legs, mandibles and antennae fuscous, palpi leoninous.

Head: Two-thirds of width of pronotum. Outline campanulate. Clypeus wide, distinctly convex anteriorly, fused with clypeal wings, not separated from supra-antennal plates, with raised sub-square field at middle, separated from frons by broad flattened shiny transverse furrow. Frons with two raised paramedian carinae, joining anteriorly into a V-like keel, with two small glossy teeth bilaterally anterior to keel, with two short parallel carinae paralaterally at base; frons separated from supraorbital plates by flattened broad area; with conspicuously deep and broad pit at front-eye level; supra-antennal and supraorbital plates margined, margin of supraorbital plate distinctly raised, carina-like, supra-antennal plates slightly vaulted. Base with moderately broad emargination at middle, laterally obtuse-angled (angle 103°-105°), with small but distinct tooth at angle, with small notch anterior tooth. Eyes large, convex, genae slightly convex, both of them just visible from above, with long ovoid shape and straight posterior margin in lateral view. Antenna with segments five to ten elongate (L/W 1.3), densely pubescent, segments two to four scarcely pubescent, scapus with longitudinal reticulation. Labrum convex anteriorly. Mandible of moderate size, regularly arcuate, more distinct at apex. Mentum small, pentagon-like shaped, isodiametrically reticulated, without tooth, epilobes wide, projecting and angled anteriorly, margined completely, surface with medium sized pits, shiny.

Pronotum (Fig. 38 View Figures 36–43 ): Outline rectangular, transverse, a third wider than long. Lateral margin regularly convex, maximum width at end of second third. Lateral margin distinctly crenulated, with 10(-12) distinct tubercles, with a notch at posterior angles. Base oblique laterally, with declining flat ridge at middle. Disc flattened at middle in lateral view, with two distinctly raised paramedian carinae running parallel to median line and diverging at base, with row of ten small tubercles arranged saw-like, with median line medium sized, ending in broad pits anteriorly and posteriorly; with two additional shorter carinae bilaterally at base, joining with the paramedian carinae and forming tooth-like tubercle at base pointing posteriorly, with long anterior extension, with short inner lateral carina, with indistinct outer lateral carinae. All carinae sub-crenulate. Lateral margin broadly wing-like bent up, with six large and deep transverse pits. Space between carinae and pits smooth at middle.

Elytron: Flattened in anterior half (lateral view), convex in frontal view. Sub-elongate, slightly convex laterally, maximum width at middle. Pseudohumerus sub-rectangular (angle 96°), with laterally projecting tooth. Apex rounded, without tooth at suture. Disc with interneur six crenulated; interneur three slightly carinate in basal sixth, interneur two running up to apex as slightly convex line, conspicuously raised, reaching apex; interneur four running in parallel to interneur six, not reaching base, distinctly shortened apically. Interneur five and six with two rows of serial pits, the latter ones distinctly merging transversally.

Hind wings: Fully developed.

Lower surface: Antennal channel of pronotum with isodiametric reticulation. Pseudoepipleura with row of small pits, lateral margin of elytron smooth. Metepisternum elongate, with broad longitudinal groove reticulated isodiametrically. Metasternum with distinct pits partly joining; abdominal sternite one with longitudinal reticulation, sternite two with isodiametric reticulation, other sternites with densely arranged pits, with isodiametric reticulation laterally. Sternum four to six sulcate. Sternum five to six with isodiametric reticulation laterally, sternum six with longitudinal large pit apically.

Legs: Profemora with surface reticulated. Protibia with short, moderately curved terminal spine, laterally with three larger and three smaller teeth, dorsally and ventrally with two carinae. First tarsomeres distinctly elongated, as long as tarsomere two to four together.

External sexual dimorphism: Last visible sternum in females with lateral and central longitudinal carinae slightly more distinct developed as in males.

Male genitalia (Fig. 64 View Figures 64–67 ): Median lobe small, in dorsal view slightly arcuate in basal half, distinctly arcuate in apical half, in lateral view distinctly cracked at beginning of apical third, with fine pili laterally in apical half, apex spatulate, in cross section with concavity at apex. Oroficium nearly half as long as median lobe. Endophallus with group of conspicuously regularly arranged microtrichia near oroficium, laterally with a small group. Dorsal paramere elongated, gently sinuate apically, with hyaline apex, with short apophyses, ventral one relatively long, both parameres slightly distorted.

Female genitalia (Fig. 79 View Figures 77–83 ): Coxostylus of moderate size, distinctly broadened at base, distinctly curved, moderately acute at apex, with indistinct carina dorsally in apical third, at end of basal third with one large and seven slender nematiform setae laterally, SSO with one fine pilus.

Variation: On the head, the lateral margin of the supra-antennal and supraorbital plates shows a different degree of the slight crenulation. In one of the paratypes the clypeal wings are separated from the supra-antennal plates by slight obtuse notches. The pronotum with the lateral margin shows an inter- and intra-specific variation of the number of tubercles (12 to 15). The protibia exhibits in some specimens one or two additional small teeth with a seta. In one specimen from "Narobi b. Tanga" the pronotum is slightly more constricted anteriorly.

Etymology.

The name refers to the location Utete in Tanzania where most of the type material was found.

Distribution

(Fig. 84 View Figure 84 ). Known from the type locality in the north-east of Tanzania. The abbreviation on the label ("D. O. Afr.") refers to the former notation Deutsch Ost Afrika (German East Africa).