Dentisphaeramaxema

Páll-Gergely, Barna, Jochum, Adrienne & Asami, Takahiro, 2017, Three New Species And A New Genus Of Hypselostomatidae (Gastropoda: Pulmonata) From Cong Troi Cave, Northern Vietnam, Acta Zoologica Academiae Scientiarum Hungaricae 63 (3), pp. 327-341 : 333-336

Dentisphaera Páll- GergelyetJochumgen. n.

Typespecies: Dentisphaeramaxema Páll- GergelyetJochum, sp. n.

Differentialdiagnosis: Dentisphaera differsfrom Angustopila bythepres- enceofsixteethintheaperture, andtheelongationofthetuba (mostcon- spicuouslyvisiblefromdorsalview).

Etymology: Thename Dentisphaera referstothepresenceofmultipleteethinthe

peristomeandtheovoid, almostsphericalshapeoftheshell.

Dentisphaeramaxema Páll- GergelyetJochumsp. n.

( Figs 1D, 4, 7C–D)

Typematerial: C04-N74 (localitycode), NorthernVietnam, HaGiangProvince, Quan BaDistrict, TamSon, CongTroiCave, 980 m, 23°02’44.7”N, 104°59’23.9”E, WGS84, internal canyon, goursofdrippingwater, leg. G. TOMASIN, 29.01.2010., HolotypeMZUFGC/49408; Samedata, MZUFGC/49409/ 12 paratypes; Samedata, HNHM 99732 / 1 paratype; Same data, PGB / 2 paratypes; Samedata, MZUFGC/49410/1 juvenileshell (notparatype); C05-N5 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, TamSon, Cong TroiCave, 980 m, 23°02’44.7”N, 104°59’23.9”E, WGS84, attheentrance, lakewithdrippingwater, leg. G. TOMASIN, 29.01.2010., 2 corrodedparatypes, MZUFGC/49397; C03-N39 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, TamSon, Cong TroiCave, 980 m, 23°02’44.7”N, 104°59’23.9”E, WGS84, attheentrance, goursofdripping water, leg. G. TOMASIN, 28.01.2010., Specimen1 (figuredparatype), additional 8 paratypes, MZUFGC/49402; Samedata, 4 shellfragments (2 apical, 2 lowerparts; notparatypes), MZUF GC/49403 .

Diagnosis: Shelltiny, ovate- conoidwithshoulderedwhorls, umbilicus narrow, tubaprotruding, sixrobustteeth (1 parietal, 1 angular, 2 palatal, 2 columellar).

Description: Shellminute, ovate, veryfragile, lightgreywithspireconvexlyconicwith obtuseapex; protoconchrecessedtotherightatantepenultimatewhorl; nuclearwhorlbe- ginswithapatternofgranulosewrinkles, mergingintoapatternofdense, shallowpits; the 4.5 convex, unevenlyincreasingwhorlsareseparatedbyadeepsuture; teleoconchsculpture finelyretiformwithraisedequidistantly- spacedspiralthreads; smallaperturepiriform; six prominent, deeplysetdenticlesformrobustbarriers; parietaldenticleslightlycontortedand aligneddirectlyoppositethedeeper- set, palataldenticle; angulardenticleslantstotheright; bodywhorlthick; apertureobliquetoshellaxis; tubaendofbodywhorldetachedandpro- jectingslightlydownwardpastpenultimatewhorl (lateralview); tubadeflectstotherightof theumbilicus; umbilicalzonefaintlyreticulate; umbilicusnarrowanddeep.

Measurements (in mm): SH = 0.87–1.06; SW = 0.85–0.96; AH = 0.38–0.45; AW = 0.38–

0.44 (n = 18) ( Table 2).

Etymology: ThespecificepithetderivesfromtheHungarianword “mákszem” (poppy seed), referring to the tiny shell. This name is to be used as a noun in apposition.

tionsseeTable 1.

Distribution: Dentisphaeramaxema sp. n. isknownfromsedimentsam-

plesfromtheCongTroiCaveinNorthernVietnam.

Fig. 5. Holotype of Tonkinospira tomasini sp. n. A–D = different views of the shell, E = proto- conch, F = sculpture of the base of the shell (umbilical side), G = sculpture of the body whorl