Zyras (Zyras) wunderlei, Assing, 2016

Assing, Volker, 2016, On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae), Beiträge Zur Entomologie = Contributions to Entomology 66 (1), pp. 13-111 : 106-107

Zyras (Zyras) wunderlei spec. nov.

( Figs 184–186 , 461–465 )

Type material: Holotype ♂: “ Indonesia, Central Bali , Pupuan, Wasserfall , 700 m, Uferschotter, 14.06.94 P. Wunderle / Holotypus ♂ Zyras wunderlei sp. n. det. V. Assing 2015” ( cAss).

Etymology: This species is dedicated to my friend and long-time field companion Paul Wunderle (Mönchengladbach), who collected the holotype.

Description: Body length 6.8 mm; length of forebody 3.3 mm. Coloration: head and pronotum dark-brown; elytra reddish-brown; abdomen dark-brown, with segments VIII–X and the posterior portion of tergite VII paler; legs pale-yellowish; antennae brown, with antennomeres I–II and the base of III reddish and antennomeres IX–XI pale-yellowish (except for the infuscate base of IX); maxillary palpi dark-yellowish with the terminal palpomere pale-yellowish.

Head ( Fig. 184 ) distinctly transverse, with rather coarse and dense punctation, except for a small patch in median portion, and with rather dense, long, erect, and dark pubescence. Eyes somewhat longer than postocular region in dorsal view. Antenna ( Fig. 185 ) 2.3 mm long; antennomeres IV approximately as long as broad, V indistinctly transverse, VI–X of increasing width and increasingly transverse, X slightly more than half as broad as long, and XI of conical shape and short, distinctly shorter than the combined length of IX and X.

Pronotum ( Fig. 184 ) 1.16 times as broad as long, and 1.22 times as broad as head, rather weakly convex in crosssection; lateral margins weakly sinuate in posterior half (dorsal view); punctation dense, moderately coarse, and rather shallow; pubescence dense, long, erect, and dark.

Elytra ( Fig. 184 ) 0.78 times as long as pronotum; punctation defined, very dense, and moderately coarse; interstices much narrower than diameter of punctures; pubescence dense, long, erect, and pale-brown. Hind wings fully developed. Metatarsomere I nearly as long as the combined length of II–IV.

Abdomen ( Fig. 186 ) broad, slightly broader than elytra and with moderately deep anterior impressions on tergites III–V; all sternites with dense and long, semierect to erect brown pubescence; anterior impressions of tergites III–V each with a transverse series of sparse and fine non-setiferous punctures; tergites III–IV with numerous setiferous punctures bearing long and semierect dark setae at posterior margin and in lateral portions, with scattered setiferous punctures on disc, and with interspersed micropunctation; tergite V with setiferous punctures bearing long and semi-erect dark setae at posterior margin and on disc, and with interspersed micropunctation; tergite VI with a transverse band of numerous and moderately coarse non-setiferous punctures anteriorly, and with setiferous punctures bearing long and semi-erect setae in posterior two-thirds; tergite VII with a broad transverse band of numerous and moderately coarse non-setiferous punctures in anterior third and with setiferous punctures not arranged in distinct series in posterior two-thirds; tergite VIII with numerous dark long setae in posterior two-fifths; posterior margin of tergite VII with palisade fringe; tergite VIII ( Fig. 464 ) with dense long pubescences posteriorly, posterior margin with small and shallow median concavity.

♂: sternite VIII ( Fig. 465 ) much longer than tergite VIII, with dense and long pubescence in posterior half, its posterior margin convex; median lobe of aedeagus ( Figs 461–462 ) 1.0 mm long; ventral process rather slender and apically acute; paramere ( Fig. 463 ) approximately as long as median lobe and with very short and apically obliquely truncate apical lobe.

Comparative notes and comment: As can be inferred from external (robust body; broad abdomen; dense punctation with long and erect pubescence) and the male sexual characters (apical lobe of paramere very short), Z. wunderlei belongs to the group of species allied to Z. hirtus (KRAATZ, 1859) , which, aside from Z. hirtus , includes several named and unnamed species from the southern East Palaearctic and the Oriental regions, such as Z. setosivestis SCHEERPELTZ, 1965 ( Myanmar; male unknown), Z. funestus ( DVOŘÁK, 1996) ( Vietnam; male unknown), and Z. flexus ASSING, 2016 ( China) . In size and habitus, Z. wunderlei is similar to Z. funestus and Z. flexus , but distinguished as follows:

– from Z. funestus by much paler legs ( Z. funestus : apical halves of femora dark-brown), paler apical antennomeres IX–XI ( Z. funestus : antennomeres IX–X dark-brown, XI reddish), longer and less strongly incrassate antennae with less transverse antennomeres V–X ( Z. funestus : antenna approximately 2.0 mm long), a larger and more transverse pronotum ( Z. funestus : pronotum 1.1 times as broad as long), and by the distinctly finer punctation of the abdominal tergites VI and VII;

– from Z. flexus bymuchpalerlegs ( Z. flexus : apicalhalves of femora dark-brown), the coloration of the antennae ( Z. flexus : antennomeres I–X black, XI yellow), coarser and denser punctation of the head, the median concavity of the posterior margin of tergite VIII (absent in Z. flexus ), and by the distinctly larger aedeagus with an apically less strongly bent ventral process. For illustrations of Z. funestus and Z. flexus see ASSING (2015d) and ASSING (2016), respectively.

Distribution and natural history: The type locality is situated in central Bali, Indonesia. The holotype was collected from scree near a waterfall.

Figures

Figs 181–196. Zyrasporrectus(181–183),Z.wunderlei(184–186),Pedinopleurusnotabilis (187–189),andAenictoides derivatus (190– 196): forebody (181, 184, 187, 190); antenna (182, 185, 188, 192); abdomen (183, 186, 194); tergites IX–X (189); forebody in lateral view (191); thorax in ventral view (193); abdomen in lateral view (195); abdomen in ventral view (196). Scale bars: 181–188, 190–196: 1.0 mm; 189: 0.5 mm.

Figs 459–471. Zyras porrectus (459–460), Z. wunderlei (461–465), and Aenictoides derivata (466–471): median lobe of aedeagus in lateral and in ventral view (459–462, 468–469); paramere (463, 470); male tergite VIII (464); male sternite VIII (465); maxilla (466); labium (467); apical lobe of paramere (471). Scale bars: 459–466, 468–470: 0.2 mm; 467, 471: 0.1 mm.