Zaiwa pankowskiorum, LYUBARSKY & TIHELKA & CAI & PERKOVSKY, 2021

LYUBARSKY, GEORGY YU., TIHELKA, ERIK, CAI, CHEN-YANG & PERKOVSKY, EVGENY E., 2021, Zaiwa: A new genus of the family Lophocateridae from mid-Cretaceous Burmese amber (Coleoptera: Cleroidea), Palaeoentomology 4 (3), pp. 218-222 : 219-221

Zaiwa pankowskiorum sp. nov.

( Figs 1–3 )

Type material. Holotype (male): SIZK Bu-300 , Burmese amber . Syninclusions: two Cecidomyiidae , Scelionidae, Brachycera.

Etymology. The species is named after Madeline Pankowski and her father Mark Pankowski (Rockville, Maryland, USA) for their efforts to find, purchase, and donate important fossils to museums where they can be studied. They donated the holotype of this species to the amber collection of the Schmalhausen Institute of Zoology, the National Academy of Sciences of Ukraine.

Diagnosis. As for the genus.

Locality and horizon. Amber mine in the Hukawng Valley, Myitkyina District, Kachin State, Myanmar; Albian/ Cenomanian boundary, mid-Cretaceous.

Description. Body length 2 mm, width 0.8 mm, elongate, rather convex dorsally and flattened ventrally ( Fig.1 ). Entire body light brown; dorsum with uniformly long and dense suberect hairs, particularly at elytral margins ( Fig. 2C ). Dorsum with subuniform, rather large, suboval punctures, spaces between them more than the diameter of one puncture. Ventral side with fine and sparse punctation.

Head dorsally with coarse punctures. Frontoclypeal suture present ( Fig. 2A ). Mandibles appearing moderately developed, gradually curved apically. Maxillary palpi apparently 4-segmented, distalmost segment suboval, width at apex almost half the maximum segment length. Distalmost segment of maxillary palpus with two large papillae at the apex, more than twice as long as thick. Palpomere 3 about 1.7 times as long as wide. Labial palpi 3-segmented, apical segment nearly oval. Eyes rather large, coarsely facetted, globular, distance between them about 4 times the eye transverse diameter ( Fig. 2A ). Antennae moderately long and reaching the posterior edge of pronotum ( Fig. 2B ); antennomere 1 (scape) rather large and oval; antennomere 2 large and oval, 1.6 times shorter than scape and 1.3 times as long as wide; antennomere 3 narrow, as long as second antennomere; antennomere 4 1.5 times as long as wide; antennomeres 5–6 nearly as long as wide; antennomere 7 is shortest, at least 1.4 times as wide as long; antennomeres 8– 10 forming a loose and slightly dorsoventrally compressed club that is markedly thicker than scape and only 1.2 times shorter than antennomeres 1–7 taken together; ultimate antennomere 2.3 times as long as scape and 2.2 times as long as antennomeres 8 and 9 taken together.

Pronotum transverse, about 1.8 times as wide as long with subexplanate sides, moderately convex at disk, widest behind the middle. Pronotal margins evenly arcuate, moderately crenulate. Anterior and posterior angles rounded, not projecting. Scutellum subquadrate, transverse, 1.3 times as wide as long ( Fig. 2C ).

Prosternumgentlyandnarrowlyconvexalongthemiddle with a rather long and narrow intercoxal process. Prosternal process narrow, 3.5 times thinner than procoxa width, lacking a carina ( Fig. 2B ). Procoxae distinctly transverse. Procoxal cavities widely open posteriorly. Mesoventrite moderately short, apparently of usual structure.Mesocoxae subtriangularoval, narrowly separated, minimum distance approximately the same as between the anterior coxae. Mesoventral process flat. Metacoxae transverse, narrowly separated. Metaventrite slightly convex and with median excavation at its posterior edge between metacoxae. Metepisterna subtriangular and widened anteriorly.

Legs moderately long. Tibiae thin, about as long as femora and slightly widening apically, with row of distinct spines along outer margin (nine in protibiae). Tarsal formula ( Fig. 2D ) 5-5-5, simple, tarsomere 5 distinctly longer than tarsomeres 2–4 combined, apex twice as wide as its base. Claws simple, slightly swollen at base.

Elytra complete, elongate, 1.5 times as long as their combined width, nearly 3 times as long as pronotum, widest in middle, subparallel-sided, apices independently rounded. Elytral surface with moderately long, suberect hairs ( Fig. 2C ), surface with closely spaced and more or less regular rows of punctures. Epipleura at anterior third of elytra not wide (nearly 1.5 times narrower than scapus) and gradually narrowing and becoming obsolete at apex.

Abdomen with 5 ventrites; ventrite 1 apparently longest, ventrite 2 somewhat shorter, ventrites 3–5 subequal.

Figures

FIGURE 1. Zaiwa pankowskiorum gen. et sp. nov. in mid-Cretaceous amber from northern Myanmar. A, Dorsal view. B, Ventral view. C, Lateral view.

FIGURE 2. Zaiwa pankowskiorum gen. et sp. nov. in mid-Cretaceous amber from northern Myanmar. A, Head and pronotum, dorsal view. B, Head and prothorax, ventral view. C, Anterior part of elytra, showing dense setae. D, Hind tarsi.

FIGURE 3. Line drawing of Zaiwa pankowskiorum gen. et sp. nov. in mid-Cretaceous amber from northern Myanmar.