Vexillum pseudoschafferi Biskupič, 2020

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 53-54

Vexillum pseudoschafferi Biskupič, 2020

Figs 14E 1 –E 2 , F 1 –F 2 , G

* Vexillum pseudoschafferi sp. nov. — Biskupič 2020: 83, figs. 6–7.

Type material. Holotype: Z 40033, SL: 11.7 mm, MD: 4.4 mm, Rohožník–Konopiská ( Slovakia), illustrated in Biskupič (2020, figs 6A–B), 14E 1 –E 2 . Paratypes: Z 40034, SL: 6.8 mm, MD: 2.7 mm, Rohožník–Konopiská (Slovakia), figs 14F 1 –F 2 ; Z 40035, SL: 6.3 mm, MD: 2.6 mm, Rohožník–Konopiská ( Slovakia), fig 14 G. All types stored in the Natural History Museum of Slovak National Museum , Bratislava, Slovakia .

Revised description. Shell medium sized with slightly scalate spire. Protoconch high conical of three smooth whorls. Teleoconch of seven subcylindrical whorls. Sculpture of prominent, convex, orthocline to weakly opisthocline axial ribs (up to 20 on penultimate whorl), separated by interspaces of about equal width. Interspaces with spiral cords, separated by narrow grooves, not crossing axial ribs. Weakly nodulose subsutural cord. Last whorl rapidly contracting with deep basal concavity. Rounded spiral cords on base and fasciole. Aperture moderately wide. Columellar callus indistinct. Columella with four prominent folds. Outer lip lirate. Siphonal canal moderately short, straight, with shallow siphonal notch.

Discussion. This species is reminiscent of the extant Indo-West Pacific Vexillum micra Pilsbry, 1921 , which differs mainly in its subhorizontal columellar folds and more prominent spiral cords.

Vexillum pseudoschafferi differs from Pusia schafferi ( Meznerics, 1933) in its sculpture, but differs in its stout shape, scalate spire, and the presence of a subsutural cord.

Palaeoenvironment. Shallow marine environments in corallinacean marls ( Biskupič 2020).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Rohožník–Konopiská (Slovakia) ( Biskupič 2020).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 14 A1–A2. Pusia paraleucozona (Boettger, 1906), Drnovice u Vyškova (Czech Republic), NHMW 1865/0036/0010. B1–B2. Pusia paraleucozona (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360345. C1–C2. Pusia szobbiensis (Halaváts, 1884), lectotype, Inv. nr. M107, Szob (Hungary). D. Vexillum szobbiensis (Halaváts, 1884), paralectotype, Inv. nr. M107, Szob (Hungary). E1–E2. Vexillum pseudoschafferi Biskupič, 2020, holotype, Z 40033, Rohožník–Konopiská (Slovakia). E1–E2. Vexillum pseudoschafferi Biskupič, 2020, paratype, Z 40034, Rohožník–Konopiská (Slovakia). G. Vexillum pseudoschafferi Biskupič, 2020, paratype, Z 40035, Rohožník–Konopiská (Slovakia).