UNGULINIDAE J. E. Gray, 1854

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 777-779

Family UNGULINIDAE J. E. Gray, 1854

adansoni, Cyclina –Deshayes, 1853a: 32. — Type locality: Senegal. — Type age: Recent. This was proposed by Deshayes in his Veneridae catalogue, based on Adanson’s unavailable “chama felan”.The first name available for this species was Venus diaphana Gmelin, 1791 . — Current status: Diplodonta diaphana (Gmelin, 1791) and the Deshayes species belongs in its synonymy ( Cosel & Gofas 2019: 828-829, fig. 16.25).

aizyensis , Diplodonta – Deshayes, 1857-DescrCoqFoss: 47, pl. 47, figs 13-16; 1858-DescrCoqFoss: 612-613. — Type localities: Aizy-Jouy & Coeuvres-et-Valsery. — Type age: Eocene (Ypresian). Diplodonta (Zemysina) aizyensis Deshayes, 1857 ( Glibert & Van de Poel 1967: 7). — Current status: Phlyctiderma aizyense (Deshayes, 1857) ( Le Renard & Pacaud 1995: 75) .

auversiensis , Diplodonta – Deshayes, 1857-DescrCoqFoss: 46, pl. 46, figs 20-22; 1858-DescrCoqFoss: 617-618. — Type localities:Auvers-sur-Oise & Valmondois. — Type age: Eocene (Bartonian). — Current status: Diplodonta (Diplodonta) auversiensis Deshayes, 1857 ( Le Renard & Pacaud 1995: 75).

bidens , Diplodonta – Deshayes, 1857-DescrCoqFoss: 42, pl. 42, figs 17-19, 45, pl. 45, figs 28-30; 1858-DescrCoqFoss: 619-620. — Type localities: Hermonville, Damery, Fleury-la-Rivière, Le Fayel & Caumont. — Type age: Eocene (Lutetian-Bartonian). — Current status: Diplodonta (Zemysia) bidens Deshayes, 1857 ( Glibert & Van de Poel 1967: 11; Le Renard & Pacaud 1995: 76; Pacaud 2008: 87).

biimpressa , Diplodonta – Deshayes, 1857-DescrCoqFoss: 45, pl. 45, figs 22-24; 1858-DescrCoqFoss: 618. — Type localities: Cuise-la-Motte & Retheuil. — Type age: Eocene (Ypresian). — Current status: Felaniella (Felaniella) biimpressa (Deshayes, 1857) ( Glibert & Van de Poel 1967: 9; Le Renard & Pacaud 1995: 76).

brochii , Lucina – Deshayes, 1850-TraitElem: 786. New species based on Lucina globosa (Gmelin, 1791) , of Brocchi, 1814 ( Brocchi 1814: 553-554), non (Gmelin, 1791) [ Venus ]. — Type locality: Andona, Italy. — Type age: not stated. — Type material: Museo Civico di Storia Naturale di Milano (MCSN) 442, lectotype, Brocchi’s specimen. Name should be corrected to L. brocchii because it was clearly misspelled, having been named for Brocchi (ICZN Code Art. 32.5). Diplodonta (Zemysina) brochii (Deshayes, 1850) ( Glibert & Van de Poel 1967: 7) . — Current status: Diplodonta brocchii (Deshayes, 1850) ( Carrozza 1983: 141-144, fig. 1, lectotype; Sabelli et al. 1990: 302; Cossignani & Ardovini 2011: 53, 440; Janssen & Krylova 2014: 76; Huber 2015: 329; Cosel & Gofas 2019: 824-825; Alf & Haszprunar, in Alf et al. 2020: 390-391, pl. 382).

coelata , Diplodonta – Deshayes, 1857-DescrCoqFoss: 47, pl. 47, fig. 24; 1858-DescrCoqFoss: 614-615. — Type locality: Noailles. — Type age: Paleocene (Thanetian). — Current status: Diplodonta (Diplodonta) coelata Deshayes, 1857 ( Glibert & Van de Poel 1967: 5; Le Renard & Pacaud 1995: 75).

consors , Diplodonta – Deshayes, 1857-DescrCoqFoss: 46, pl. 46, figs 13-16; 1858-DescrCoqFoss: 622-623. — Type locality: Cuise-la-Motte. — Type age: Eocene (Ypresian). Felaniella (Bruetia) consors (Deshayes, 1857) ( Glibert & Van de Poel 1967: 11-12) . — Current status: Bruetia consors (Deshayes, 1857) ( Le Renard & Pacaud 1995: 76; Pacaud 2008: 87). decipiens , Diplodonta – Deshayes, 1857-DescrCoqFoss: 46, pl. 46, figs 27-29; 1858-DescrCoqFoss: 615-616. — Type localities: Thiverval-Grignon, Parnes & Damery. — Type age: Eocene (Lutetian). Deshayes noted that he had figured this species as Lucina renulata Lamarck, 1806 , in his first Paris Basin edition (1825-DescrCoqFoss: 93-94). Pezant (1911: 110) proposed Diplodonta depulsa as replacement name for “ Diplodonta renulata Desh. non Lam.”, which would make it a synonym of Diplodonta decipiens Deshayes. — Current status: Diplodonta (Zemysina) decipiens Deshayes, 1857 ( Glibert & Van de Poel 1967: 7; Le Renard & Pacaud 1995: 75; Pacaud 2008: 87).

duplicata , Diplodonta – Deshayes, 1857-DescrCoqFoss: 45, pl. 45, figs 25-27; 1858-DescrCoqFoss: 619. — Type localities: Abbecourt & Noailles. — Type age: Paleocene (Thanetian). — Current status: Felaniella (Zemysia) duplicata (Deshayes, 1857) ( Glibert & Van de Poel 1967: 11; Le Renard & Pacaud 1995: 76).

eudora , Diplodonta – Deshayes, 1857-DescrCoqFoss: 46, as D. eurorae , pl. 46, figs 4-6; 1858-DescrCoqFoss: 621. — Type localities: Aizy-Jouy, Hérouval & Mercin-et-Vaux. — Type age: Eocene (Ypresian).— Current status: Felaniella (Felaniella) eudora (Deshayes, 1857) ( Glibert & Van de Poel 1967: 10; Le Renard & Pacaud 1995: 76).

fragilis , Diplodonta – Deshayes, 1857-DescrCoqFoss: 46, pl. 46, figs 7-9; 1858-DescrCoqFoss: 623, non Braun, 1850. — Type localities: Châlons-sur-Vesle & Jonchery-sur-Vesle. — Type age: Paleocene (Thanetian). Diplodonta catalaunensis Cossmann & Lambert, 1884 ( Cossmann & Lambert 1884: 87), unnecessary replacement name because there is an available synonym. — Current status: synonym of Bruetia inaequalis (Deshayes, 1857) (Pacaud herein).

grata , Diplodonta – Deshayes, 1857-DescrCoqFoss: 47, pl. 47, figs 3-5; 1858-DescrCoqFoss: 611-612. — Type localities: Mouchy-le-Châtel, Les Groux & Parnes. — Type age: Eocene (Lutetian). — Current status: Diplodonta ( Zemysina ?) grata Deshayes, 1857 ( Glibert & Van de Poel 1967: 8). Phlyctiderma gratum (Deshayes, 1857) ( Le Renard & Pacaud 1995: 75) .

grignonensis , Diplodonta – Deshayes, 1857-DescrCoqFoss: 47, pl. 47, figs 9-12; 1858-DescrCoqFoss: 616-617. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). Diplodonta grignonensis Deshayes, 1857 ( Cossmann & Peyrot, 1912b: 244). Timothynus? grignonensis (Deshayes, 1857) (Chavan, 1962: 16) . — Current status: synonym of Diplodonta (Zemysina) decipiens Deshayes, 1857 ( Glibert & Van de Poel 1967: 7; Le Renard & Pacaud 1995: 75).

guyerdeti, Diplodonta – Deshayes, 1866b: 330, 332, 336-337, pl. 7, fig. 2. — Type locality: Montmartre. — Type age: Eocene (Bartonian). — Current status: Diplodonta (? Zemysina ) guyerdeti Deshayes, 1866 ( Le Renard & Pacaud 1995: 75).

herouvallensis , Goodallia – Deshayes, 1858-DescrCoqFoss: 63, pl. 63, figs 27-29 [fig. 29 not cited in text]; 1860-DescrCoqFoss: 786; 1865-DescrCoqFoss: 665 [error noted]. — Type locality: Hérouval. — Type age: Eocene (Ypresian). — Type species (SD Dall, 1903) of Microstagon Cossmann, 1896 . Microstagon herouvalense [sic] (Deshayes, 1858) ( Glibert & Van de Poel 1970: 1967: 4). — Current status: Microstagon herouvallense (Deshayes, 1858) ( Le Renard & Pacaud 1995: 80) .

inaequalis , Diplodonta –Deshayes, 1857-DescrCoqFoss: 46, pl. 46, figs 10-12; 1858-DescrCoqFoss: 623-624. — Type localities: Châlons-sur-Vesle & Jonchery-sur-Vesle. — Type age: Paleocene (Thanetian). 1858: 552. Felaniella (Bruetia) inaequalis (Deshayes, 1857) ( Glibert & Van de Poel 1967: 12) . — Current status: Bruetia inaequalis (Deshayes, 1857) ( Le Renard & Pacaud 1995: 76) . incrassata , Goodallia – Deshayes, 1858-DescrCoqFoss: 59, pl. 59, figs 18-20; 1860-DescrCoqFoss: 785. — Type locality: Hérouval. — Type age: Eocene (Ypresian). — Current status: Microstagon laevigatum [Deshayes, 1858] incrassatum (Deshayes, 1858) ( Le Renard & Pacaud 1995: 80).

ingens , Diplodonta – Deshayes, 1857-DescrCoqFoss: 47, pl. 47, figs 1-2; 1858-DescrCoqFoss: 611. — Type locality: Chalons-sur-Vesle. — Type age: Paleocene (Thanetian). — Current status: Diplodonta (Zemysina) ingens Deshayes, 1857 ( Glibert & Van de Poel 1967: 8; Le Renard & Pacaud 1995: 75; Leroy et al. 2014: 28, pl. 11, fig. 5a-b).

laevigata , Goodallia – Deshayes, 1858-DescrCoqFoss: 63, pl. 63, figs 24-26; 1860-DescrCoqFoss: 784. — Type localities: Parnes, Chaumont-en-Vexin & Les Groux. — Type age: Eocene (Lutetian). — Current status: Microstagon laevigatum (Deshayes, 1858) ( Glibert & Van de Poel 1967: 4-5; Le Renard & Pacaud 1995: 80).

laevigata , Lucina – Deshayes, 1825-DescrCoqFoss: 94, 7, pl. 15, figs 9-10; 1832-EncyMeth: 374; 1858-DescrCoqFoss: 620, as Diplodonta . — Type locality: Abbecourt. — Type age: Paleocene (Thanetian). — Current status: Felaniella (Felaniella) laevigata (Deshayes, 1825) ( Glibert & Van de Poel 1967: 10; Le Renard & Pacaud 1995: 76; Leroy et al. 2014: 28, pl. 11, fig. 4a-b).

lamberti , Diplodonta – Deshayes, 1857-DescrCoqFoss: 47, pl. 47, figs 6-8; 1858-DescrCoqFoss: 616. — Type localities: Mercin-et-Vaux, Cuise-la-Motte & Hérouval. — Type age: Eocene (Ypresian). — Current status: Diplodonta (Zemysina) lamberti (Deshayes, 1857) ( Glibert & Van de Poel 1967: 8; Le Renard & Pacaud 1995: 75).

lenticularis , Cyrenella – Deshayes, 1855c: 341. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 20110007, 3 probable syntypes. — Current status: taxon inquirendum ( Huber 2015: chapter 5 on CD).

lucinoides , Venus – Deshayes, 1825-DescrCoqFoss: 146-147, 12, pl. 23, figs 12-13; 1832-EncyMeth: 1123; 1835-HistNatAnim2: 379; 1839-HistNatAnim3: 620; 1850-TraitElem: 818; 1857-TraitElem: iv, pl. 14 bis, figs 10-12, as Cyrenella ; 1858-DescrCoqFoss: 612, as Diplodonta . — Type locality: La Chapelle-en-Serval. — Type age: Eocene (Bartonian). — Current status: Phlyctiderma lucinoides (Deshayes, 1825) ( Le Renard & Pacaud 1995: 75) .

moretonensis , Cyrenella – Deshayes, 1855c: 341. — Type locality: Moreton Bay, Queensland, Australia; Cuming coll. — Type age: Recent. — Type material: NHMUK 201105, syntype. Joannisiella moretonensis (Deshayes, 1855) ( Hedley 1913: 267, pl. 16, figs 11- 15). — Current status: synonym of Joannisiella cumingii (Hanley, 1846) [ Cyrenoidea ] ( Huber 2015: chapter 5 on CD).

oblonga , Cyrenella – Deshayes, 1855c: 341, ex Sowerby ms. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type species (OD) of Joannisia Dall, 1895 , non Monterosato, 1888; Joannisiella Dall, 1895 , replacement name. — Current status: synonym of Joannisiella oblonga (Hanley, 1846) [ Cyrenoidea ] ( Huber 2015: chapter 5 on CD).

philippinarum , Cyrenella – Deshayes, 1855c: 340, ex Sowerby ms. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1963550, 3 probable syntypes. — Current status: Joannisella philippinarum (Deshayes, 1855) ( Huber 2015: 342) .

pisiformis , Cyrenella – Deshayes, 1855c: 341. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 20110006, 1 syntype. — Current status: Zemysina pisiformis (Deshayes, 1855) ( Huber 2015: chapter 5 on CD). profunda , Lucina –Deshayes, 1850-TraitElem: 782.Bordeaux.— Type age: Miocene (Burdigalian). New species based on Lucina renulata Lamarck, 1806 , of Basterot (1825: 88), non Lamarck, 1806. — Type material: MNHN.F.A13827, syntypes. Preoccupies Lucina profunda Dufour, 1881 , which was renamed Gibbolucina aremorica Pacaud, 2020 ( Pacaud 2020c: 36). — Current status: Diplodonta profunda (Deshayes, 1850) ( Pacaud 2020c: 37) .

profunda , Diplodonta – Deshayes, 1857-DescrCoqFoss: 46, pl. 46, figs 30-33; 1858-DescrCoqFoss: 617, non Diplodonta profunda (Deshayes, 1850) [ Lucina ]. — Type locality: Mouchy-le-Châtel. — Type age: Eocene (Lutetian). — Current status: Diplodonta (Diplodonta) profunda Deshayes, 1857 ( Le Renard & Pacaud 1995: 75; Pacaud 2008: 87). Diplodonta oesia Pacaud, 2020 , replacement name ( Pacaud 2020c: 38).

punctatissima, Diplodonta –Deshayes, 1857-DescrCoqFoss: 47, pl. 47, figs 17-20; 1858-DescrCoqFoss: 613. — Type localities: Vregny & Mercin-et-Vaux. — Type age: Eocene (Ypresian). — Current status: synonym of Mysia aizyensis (Deshayes,1857) ( Cossmann 1887: 18-19) . Phlyctiderma aizyense (Deshayes, 1857) ( Le Renard & Pacaud 1995: 75) .

saincenyensis , Diplodonta –Deshayes, 1857-DescrCoqFoss: 46, pl. 46, figs 17-19; 1858-DescrCoqFoss: 622. — Type locality: Sinceny. — Type age: Eocene (Ypresian). — Current status: Bruetia saincenyensis (Deshayes, 1857) ( Le Renard & Pacaud 1995: 76) .

sphaericula , Cyrenella – Deshayes,1855c: 340.—Type locality: Moreton Bay , Queensland, Australia ; Cuming coll. — Type age: Recent.— Type material: NHMUK 201104, 3 syntypes. — Current status: Zemysina sphaericula (Deshayes, 1855) ( Huber 2015: chapter 5 on CD).

striatina , Diplodonta – Deshayes, 1857-DescrCoqFoss: 47, pl. 47, figs 21-23; 1858-DescrCoqFoss: 614. — Type localities:Le Guépelle, Beauval, Acy-en-Multien & Valmondois. — Type age: Eocene (Bartonian).— Current status: Diplodonta (Zemysina) striatina Deshayes, 1857 ( Glibert & Van de Poel, 1967: 8-9; Le Renard & Pacaud 1995: 75).

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gibbosula , Cycladicama – This was figured by Poppe (2011: 158, pl. 1074, figs 3-4) as “Deshayes, 1854”. The genus Cycladicama Valenciennes, 1854 , is regarded as a nomen dubium in the Ungulinidae , and species previously placed in it are now allocated to Joannisiella Dall, 1895 . However, there is no such species name in this genus and perhaps Joannisiella philippinarum (Deshayes, 1855) [ Cyrenella ] was intended, or perhaps Cyrenella oblonga Deshayes, 1855 , a synonym of Joannisiella oblonga (Hanley, 1846) [ Cyrenoidea ].

indica , Diplodonta – Listed by Comber (1906: 214) and by Melvill & Standen (1907: 816), as by Deshayes, then repeated in subsequent literature. It perhaps was originally copied from a label. In both cases, a nomen nudum.

radians , Diplodonta – Deshayes, 1857-DescrCoqFoss: 45, pl. 45, figs 31-33; 1858-DescrCoqFoss: 621-622. Made available byMelleville (1843: 80-81 [34, 82]. pl. 1, figs 13-14), as Lucina radians , non Conrad, 1842. — Type locality: “Tout le Laonnois”. — Type age: Eocene (Ypresian). — Current status: Lucina subradians d’Orbigny (1850b: 305, no. 156), replacement name, now Bruetia subradians (d’Orbigny, 1850) .

rotundata , Lucina – Deshayes, 1839: 11, pl. 16, figs 12-14. Sherborn (1930: 5566) listed this as a Deshayes species. However, it is clear from the Traité (1850: 785, 793) that the Deshayes use of his name was only intended to indicate a transfer of Tellina rotundata Montagu, 1803 , to Lucina , a species now listed as the European Diplodonta rotundata (Montagu, 1803) , the senior synonym of the type species of Diplodonta Bronn, 1831 (figured in Cossignani & Ardovini 2011: 53, 440; Alf & Haszprunar, in Alf et al. 2020: 391, pl. 328).