Tylopus grandis, Likhitrakarn, Natdanai, Golovatch, Sergei I., Prateepasen, Rujiporn & Panha, Somsak, 2010

Likhitrakarn, Natdanai, Golovatch, Sergei I., Prateepasen, Rujiporn & Panha, Somsak, 2010, Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae), ZooKeys 72, pp. 23-68 : 27-28

Tylopus grandis sp. n. Figs 46

Holotype

♂ (CUMZ), Thailand, Mae Hong Son Province, Pangmapha District, near Cave Pha Mon, 19.07.2008, leg. S. Panha, J. Sutcharit & N. Likhitrakarn.

Paratypes:

1 ♂, 2 ♀ (CUMZ), same lcality, together with holotype. 1 ♂ (CUMZ), same District, Mae Lana crossroads, 19.07.2008, leg. S. Panha, J. Sutcharit & N. Likhitrakarn.

Name:

To emphasize the large size of this species

Diagnosis:

Differs from congeners in the large size, coupled with a short spiniform process h, a basally only poorly delimited lobe l, and a small lobiform process z of the gonopod.

Description:

Length 41 mm (holotype), 40-42 mm (♂), 38-39 mm (♀), width of midbody pro- and metazona 3.0 and 4.5 mm (holotype), 2.8-3.0 and 4.3-4.5 mm (♂), 3.6-3.8 and 4.7-5.0 mm (♀), respectively. Coloration in alcohol dark brown to black-brown (Fig. 4 A–G): calluses, venter and antennomeres 1-5 slightly to considerably lighter, brown to light yellow-brown (Fig. 1 A–G), antennomeres 6 and 7 dark brown.

All characters as in Tylopus bispinosus sp. n., except as follows.

Antennae short and slender (Fig. 4B), reaching behind segment 3 (♂) or 2 (♀) dorsally. In width, head <collum = segments 3-4 <2 = 5-16 (♂), or head <segments 3-4 <collum <segment 2 <5-16 (♀); thereafter body gradually and gently tapering towards telson (Fig. 4 A–F).

Tegument generally rather smooth and either dull (only in places modestly shining) or shining (Fig. 4 A–G). Paraterga strongly developed (Fig. 4 A–G), lying high (at 1/4-1/5 midbody height), subhorizontal to slightly upturned laterally (Fig. 4 A–F). Transverse sulcus either absent or poorly developed, then not reaching bases of paraterga 4, always evident and reaching bases of paraterga 5-18, rather faintly rugulose at bottom. Stricture between pro- and metazona rather faintly beaded to striolate (Fig. 4 A–C). Epiproct tip evidently emarginate, pre-apical papillae very distinct (Fig. 4F, G). Hypoproct semi-circular, caudal setae strongly separated (Fig. 4G). Pleurosternal carinae visible on segments 2-15(16) (♂) or segments 2-6 (♀), mostly as low bulges anteriorly and a more or less distinct denticle posteriorly (Fig. 4B, D).

Sterna moderately setose, without modifications; a slightly notched sternal lobe between ♂ coxae 4 (Fig. 4H, I). Legs long and slender (Fig. 1B, C, H), especially so in ♂ compared to ♀ (1.7-1.8 versus 1.5-1.6 times as long as midbody height); ♂ prefemora distinctly bulged laterally (Fig. 4J), acropodites with particularly dense, nearly adpressed setae, but tarsal brushes missing.

Gonopods (Figs 5, 6) with lobe l poorly demarcated at base; spine h very small; process z not spiniform, but like a short lobe.