Triphysa nervosa biocellata Staudinger, 1901

Dubatolov, Vladimir V., Korb, Stanislav K. & Yakovlev, Roman V., 2023, A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae), Ecologica Montenegrina 61, pp. 88-123 : 116

Triphysa nervosa biocellata Staudinger, 1901

Figs. 4i–t , 6a , 16 , 30

Triphysa phryne biocellata Staudinger, 1901 , in: Staudinger, Rebel, 1901; Cat. Lep. Pal. Faunengeb. 1: 67.

Type locality: “ Amdo” (stated erroneously). By lectotype designation (designated here) the type locality is: “ Tetang mont.” [Qinghai, China]. “... a praec. var. [Dohrnii] differt al. ant. supra ocellis duobus magnis caecis (nigr.) plerumque etiam in ♂ perspicuis; al. post. subt. ocell. minoribus.”

Lectotype male, designated here, labelled: handwritten (black ink) on brown paper “ Tibet | Tetang mont1 | 90 Gr.Gr. ”; brown circle; handwritten (black ink) on white paper “Triph. | Dohrnii | Tetung- mont”; printed on pink paper “Origin.”; printed on red paper “ Lectotype ♂ | Triphysa phryne | biocellata Staudinger, 1901 | Cat. Lep. Pal. Faunengeb.: 67 | Dubatolov, Korb, Yakovlev des. 2013”.

= ssp. kintschouensis O. Bang-Haas, 1939 ( Triphyra (sic!) phryne kintschouensis O. Bang-Haas, 1939 , Dt. Ent. Z. Iris, B. 53: 53).

Lectotype male, designated here, labelled: printed on white paper “Mandschuris mer. occ | Prov. Fengtien | Kintschou | 100 m Juli”; printed on white apaper “162.”; printed on pink paper “Co- Type | O.Bang-Haas”; handwritten (by pencil) on white paper “phryne | kintschouensis”; printed on red paper “ Lectotype ♂ | Triphysa phryne | kintschouensis O.Bang-Haas, 1939 | Dt. Ent. Z. Iris, B. 53: 53 | Dubatolov, Korb, Yakovlev des. 2013”.

Diagnosis. Upperside with well developed light border. Underside with highly enlarged cubital eyespots of postdiscal row with small white centers. Female has large cubital spots often visible on the wings’ upperside. Veins on the underside are bright, the whole pattern is mottled. Pictures of genitalia published by Bozano (2002).

Material examined. 1 male (lectotype of biocellata ); 1 male (lectotype of kintschouensis ), 1 female (paralectotype of kintschouensis ) ( MHUB) ; 1 male, 1 female, Mandschuria mer. occ., Prov. Fengtien, Kintschou, 100 m, Juli (paralectotypes of kintschoensis) ; 2 males, 1 female (paralectotypes of biocellata ) ( MHUB) ; 1 male, 1 female, Tibet ( Kuku-Noor ) ( ZMHU) (paralectotypes of biocellata ) ( ZMHU) ; 1 female, Thibet, Tetung mont., [18]90, Gr. - Gr. ( MHUB) (cotype) ; 1 female, Kuku-Nor 94 Rkbl. ( MHUB) ; 2 males, 1 female, Kuku-noor ( LNK) ; 1 female, Kuku-noor ( ZFMK) ; 6 males, 6 females, Kuku-noor , Tancre ( ZSSM) ; 2 males, 2 females, Mandschuria merid., Prov. Fengtien , Kintschou, 100 m (paralectotypes of kintschouensis ) ( ZSSM) ; 1 male, 2 females, China, Prov. Chinghai, Sinig-Huangyuan , 30-V-1951, F. Eichinger ( ZSSM) ; 5 males, 5 females, Russia, Buryatia Rep., Gusinoe Ozero vill., steppe rivulet valley, 25- V-2002, leg. Shevnin ( RYB) ; 4 males, 4 females, Mongolia, Darhan , 17−27-V-1987, leg. S. Andreev ( RYB) ; 1 male, Buratia, Ulan-Ude distr., Kokorino , Gilbira river , 2-VI-[19]73 ( ISEAN) ; 5 males, 5 females, Daurian Nature Reserve,. NE coast of Zun-Torei lake, Kuku-Khadan Mt. , southern slope, 8-VI-1995, V.V. Dubatolov ( ISEAN) ; 1 male, same locality, 10-VI-1995, V.V. Dubatolov ( ISEAN) ; 1 female, same locality, 11-VI-1995, V.V. Dubatolov, R. Dudko ( ISEAN) ; 1 male, same locality, 14-VI-1995, V.V. Dubatolov ( ISEAN) ; 1 female, same locality, 7-VI-1995, V.V. Dubatolov, R. Dudko and I. Lyubechansky ( ISEAN) ; 1 female, same locality, 12-VI-1995, V.V. Dubatolov, R. Dudko ( ISEAN) ; 2 males, 2 females, Daurian Nature Reserve, NE shore of Zun-Torei lake, steppe from Batukha to Gydyrgun , 9-VI-1995, V.V. Dubatolov ( ISEAN) ; 1 female, Daurian Nature Reserve, NE shore of Zun-Torei lake, steppe between Kuku-Khadan and Gydyrgun , 9-VI-1995, V.V. Dubatolov, R. Dudko ( ISEAN) ; 1 male, 1 female, Daurian Nature Reserve, NE shore of Zun-Torei lake, near Gadyrgun Mt. , valley with no trees, 11-VI-1995, V.V. Dubatolov ( ISEAN) ; 2 males, 1 female, Daurian Nature Reserve, NE shore of Zun-Torei lake, Khadanyata valley , steppe, 11-VI-1995, V.V. Dubatolov ( ISEAN) ; 1 male, 1 female, 7 km W Barun Torei lake, southern environs of Nizhny Mukey lake near Mongolian border, steppe, 15-VI-1995, V.V. Dubatolov ( ISEAN) ; 1 male, Kiachta кбвкНМ ( ZISP) ; 1 female, cAP ( ZISP) ; 1 male, Kiachta , coll. Erschoff ( ZISP) ; 1 male, 1 female, Troizkosavsk, Mikhno , VII.[19]03 ( ZISP) ; 1 female, Ust’-Kiran on Chikoj river, east Kiachta, Khomze , 3.VI.[19]03 ( ZISP) ; 18 males, 1 female, Urga Leder ( ZISP) ; 6 males, 8 females, Sichuan, Sunnan [Songnan], 9500’, Berezovsk. [y], 15-25.V.[18]94 ( ZISP) ; 1 male, 1 female, Detung [Datung He] Gr. - Gr. кбвкНМ ( ZISP) ; 1 male, S. Tetung Mts. ( ZISP) ; 1 female, Kuku Nor coll. Avinoff ( ZISP) ; 1 female, [Nei Mongol,] Inn Shan Chingan mont. Mongolei or. 2000m Juni ( ZISP) .

Distribution. China (Sichuan (partly), Qinghai, Gansu, Nei Mongol, mountains of Western Liaonin); East Mongolia (Darkhan etc.), Russia (South Buratia (Selengan Dauria), southern part of Chita Prov.). Actually this subspecies is widely distributed: it covers all Eastern and Inner Mongolia (westwards from Darkhan) to Torei Lake Depression in the Daursky Nature reserve, as well as in Chinese provinces from Sinuan and Qinghai to Liaonin. Butterflies prefer dry steppes and perhaps semideserts.

Figures

Figure 4. Triphysa sp., imago: a – T. nervosa mongolaltaica, holotype (ZISP), upperside, b – underside; c – T. nervosa mongolaltaica, paratype, male (RYB), upperside, d – underside; e – T. nervosa mongolaltaica, paratype, female (RYB), upperside, f – underside; g – T. nervosa mongolaltaica, paratype, female (RYB), upperside, h – underside; i – T. nervosa biocellata, lectotype (MHUB), upperside, j – underside; k – T. nervosa biocellata, paralectotype, female (MHUB), upperside, l – underside; m – T. nervosa biocellata (lectotype T. phryne kintschouensis) (MHUB), upperside, n – underside; o – T. nervosa biocellata (paralectotype, female T. phryne kintschouensis) (MHUB), upperside, p – underside; q – T. nervosa biocellata, male, Russia, Buryatia Rep., Gusinoe Ozero vill., steppe rivulet valley, 25.05.2002, leg. Shevnin (RYB), upperside, r – underside; s – T. nervosa biocellata, female, Russia, Buryatia Rep., Gusinoe Ozero vill., steppe rivulet valley, 25.05.2002, leg. Shevnin (RYB), upperside, t – underside; u – T. nervosa gartoki, lectotype (MHUB), upperside, v – underside.

Figure 6. Distribution map of Triphysa species: a (above) – general map; b (below) – northern-eastern part of area of T. nervosa nervosa.

Figures 16–17. Habitats of Triphysa: 16 – T. nervosa biocellata, Central Mongolia, near Ulan-Baator (photo by A. Vaganov); 17 – T. nervosa biocellata (photo by O. Kosterin).

Figures 25–30. Male genitalia (lateral projection) of Triphysa: 25 – T. nervosa glacialis, Altai, Ukok, Mai-Pak, h–2300 m; 26 – T. nervosa tuvinica, paratype; 27 – T. nervosa arturi, paratype; 28 – T. nervosa kobdoensis, paratype; 29 – T. nervosa mongolaltaica, holotype; 30 – T. nervosa biocellata, lectotype.